Haversia Roewer, 1913

Pinto-da-Rocha, Ricardo, Benedetti, Alipio Rezende, de Vasconcelos, Eduardo Gomes & Hara, Marcos Ryotaro, 2012, New systematic assignments in Gonyleptoidea (Arachnida, Opiliones, Laniatores), ZooKeys 198, pp. 25-68 : 46

Haversia Roewer, 1913 new subfamilial assignment

Haversia Roewer, 1913: 170; Kury 2003: 129; (type species: Gonyleptes defensus Butler, 1876).

Hoggellula Roewer, 1930: 397; Kury 2003: 129; (type species: Sadocus vallentini Hogg, 1913, by monotypy). Syn. n.

Diagnosis.

Haversia resembles Pachylinae with three scutal areas, such as Corralia Roewer, 1913, Diconospelta Canals, 1934, Graphinotus Koch, 1839, Huassampilia Roewer, 1913, Neogonyleptes Roewer, 1913, Oxapampeus Roewer, 1936, Spinivunus Roewer, 1943, Tumbesia Loman, 1899, Ubatubesia B. Soares, 1945, and two species of Sadocus Sørensen, 1886 ( Sadocus conspicillatus and Sadocus polyacanthus ). Haversia can be distinguished from those genera by the unarmed and smooth (i.e. without tubercles) scutal areas and the complex apophyses of male trochanter IV: three prolateral apophyses (anterior median, posterior median and an apical apophysis).

Taxonomical note.

The two type species ( Gonyleptes defensus and Sadocus vallentini )are synonymous, resulting in a generic synonymy, and the new assignment proposed here is based on penis morphology (see note on Gonyleptes denisi ). The penis of Haversia follows the basic pattern of those gonyleptids placed in the large (and non-monophyletic) subfamily Pachylinae (ventral plate with straight anterior margin; glans without dorsal process; distal, median and basal setae of ventral plate short [seta length inferior to ventral plate width]; stylus and ventral process with variable shape). However, Haversia presents two features that stand out: (i) the penis glans projected dorsally and (ii) the detailed shape of apex of penis ventral process. Those features are typical of some Chilean genera, such as Fonckia , Neogonyleptes and Sadocus (unpublished data), and might be the evidence that those four genera are closely related.

Composition.

Monotypic.