Ancylometes concolor (Perty)

Höfer, H. & Brescovit, A. D., 2000, A revision of the Neotropical spider genus Ancylometes Bertkau (Araneae: Pisauridae), Insect Systematics & Evolution 31, pp. 323-360 : 332-356

Ancylometes concolor (Perty)

(Fig. 1-6, 28-30, 60)

Aranea unicolor: Lamarck, 1818: 102, nomen nudum (Latreille manuscript name; specimen from Brazil).

Dolomedes concolor Perty, 1833: 197 (female from Brazil; should be in ZSM but lost); Walckenaer 1837: 365 (synonym of Ctenus unicolor).

Ctenus unicolor Walckenaer, 1837: 365.

Ctenus concolor C.L. Koch, 1837: 27; 1847: 59, fig. 1458.

Ancylometes vulpes Bertkau, 1880: 114 (from Barbacena, Minas Gerais, Brazil; immature female holotype, not found in any European museum, probably lost, see Levi 1991: 203); Roewer 1954: 146; Bonnet 1955: 319; Bernarde et al. 1999: 200, fig. 3. Syn. n.

Ctenus argentinus Holmberg, 1881: 271, pl. 1, fig. 3 (from Paraná de Las Palmas, Argentina; female holotype lost). Syn. n.

Ancylometes bolivianus Tullgren, 1905: 51, pl. 7, fig. 24 (from Tatarenda, Gran Chaco, Bolivia, E. Nordenskiöld; female and immature syntypes in SMNH, examined); Schiapelli & Gerschmann de Pikelin 1970: 160, 163 (as synonym of A. bogotensis); Roewer 1954: 145; Bonnet 1955: 317. Syn. n.

Lycoctenus bahiensis Strand, 1909a: 330 (from Bahia, Brazil, Selenka; immature female holotype in ZMB, examined). Syn. n.

Lycoctenus selenkae Strand, 1909a: 331 (from Bahia, Brazil, Selenka; immature female holotype in ZMB, examined); 1909b: 427. Syn. n.

Lycoctenus paraguayensis Strand, 1909a: 335 (from San Bernardino, Paraguay, Fiebrig; male holotype no. 11432 in ZMB, examined); 1909b: 427. Syn. n.

Cupiennius argentinus Petrunkevitch, 1911: 479; Lucas 1964: 133 (as synonym of A. bogotensis).

Ancylometes paraguayensis: Petrunkevitch 1911: 540; Roewer 1954: 145; Bonnet 1955: 318.

Ancylometes bayensis: Roewer 1954: 144; Bonnet 1955: 317.

Ancylometes selenkae: Petrunkevitch 1911: 540; Roewer 1954: 145; Bonnet 1955: 319.

Ctenus paulensis Mello-Leitao, 1922: 39 (from Franca, São Paulo, Brazil; female holotype no. 8945 in MZSP, examined); Roewer 1954: 654; Bonnet 1956: 1287. Syn. n.

Cupiennius diplocellatus Mello-Leitao, 1936: 21 (from Terenos, Mato Grosso, Brazil; female holotype no. 167 should be in IBSP, lost); Roewer 1954: 657; Lucas 1964: 133(synonym of A. bogotensis). Syn. n.

Ctenus originalis Mello-Leitao, 1936: 11, fig. 22 (from Itatiaia, Rio de Janeiro, Brazil, C. Moncire; female holotype no. 42269 should be in MNRJ, probably lost); Roewer 1954: 653; Lucas 1964: 132 (as synonym of A. bogotensis). Syn. n. Ctenus iophorus Mello-Leitao, 1937: 9, fig. 10 (from Entre Rios Province, Argentina, Joerg; male and female syntypes no. 42234 should be in MNRJ, probably lost); Roewer 1954: 651; Bonnet 1956: 1283. Syn. n.

Corinoctenus anomalostomus Mello-Leitao, 1939: 140, fig. a, b (from Delta del Paraná, Buenos Aires, Argentina; female holotype no. 31823 in MACN, without epigyne, examined); Roewer 1954: 671; Carcavallo & Martinez 1960: 07 (description of male); Schiapelli & Gerschmann de Pikelin 1970: 160, 165 (as synonym of A. bogotensis). Syn. n.

Ctenus infelix Mello-Leitao, 1940: 211 (from Guaytacazes, Espírito Santo, Brazil, M. Rosa; male holotype no. 58302 in MNRJ, examined); Roewer 1954: 651. Syn. n.

Ctenus metatarsalis Mello-Leitao, 1941: 216, fig. 25 (from El Rabón, Santa Fé, Argentina, M. Birabén; female holotype no. 15194 in MLP, examined); Roewer 1954: 652; Schiapelli & Gerschmann de Pikelin 1970: 160, 165 (as synonym of A. bogotensis, erroneously citing C. guadalupei Mello-Leitao). Syn. n.

Phoneutria niveobarbata Mello-Leitao, 1945: 256, fig. 41 (from San Ignacio, Misiones, Argentina, M. Birabén; male holotype should be in MLP, probably lost); Schiapelli & Gerschmann de Pikelin 1972: 32 (as synonym of A. bogotensis). Syn. n.

Ancylometes concolor. Bonnet 1955: 317-318.

Corinoctenus greenwayi Carcavallo & Martinez, 1960: 38, fig. 1-2 (from Salta, Argentina; male holotype not found in MACN or MLP, probably lost); 1960: 8, fig. 2 (description of female); Schiapelli & Gerschmann de Pikelin 1970: 160; 165 (as synonym of A. bogotensis). Syn. n.

Synonymies. - A. vulpes Bertkau is considered a junior subjective synonym of Dolomedes concolor Perty, 1833. Specimens from the type locality (Barbacena) were examined and considered identical to the original description of A. vulpes. In cases where the type could not be examined a new synonymy is based on available figures ( Ctenus argentinus, C. originalis, C. iophorus, Corinoctenus greenwayi) or descriptions ( Cupiennius diplocellatus).

Diagnosis. - Male. Palp with median apophysis straight and broad, and relatively short. Membranous lobe on base of embolus semicircular to rectangular, not covering the base of the median apophysis (Fig. 5, 28). Female. Epigynal median plate trapezoidal, prominent with central triangular protuberance; anterior border straight or slightly invaginated (Fig. 6, 29).

Description. - Male (SMNK 1448). Prosoma dorsally dark brown with median band of yellow hairs and broad marginal bands of white hairs, both linked by diverging white stripes. Chelicerae with white hairs near base. Abdomen and legs lightbrown. Metatarsi I and II dorsally densely covered by white hairs, Metatarsi III and all tibiae irregularly spotted by white hairs. Ventral parts of body generally lighter, abdomen more reddish. Chelicerae with 3 unequal promarginal and 4 subequal retromarginal teeth.

Palp. Tibia with curved ventral and black lateral apophysis. Embolus broad at base flanked by rounded membranous lobe and describing semicircle, ending filiform beneath apex of cymbium, median part embedded in winglike conductor (Fig. 5, 28). Opposite to conductor straight hammer-like median apophysis (Fig. 5, 28).

Measurements. Total length 20.0. Prosoma 11.6 long, 10.0 wide, 3.5 high. Clypeus 0.7. Eye diameters and interdistances: AME 0.45, ALE 0.25, PME 0.5, PLE 0.45; AME-AME 0.25, AME-ALE 0.45, PME-PME 0.25, PME-PLE 0.5, AME-PLE 0.8, AME-PME 0.35, MOQ length 1.25, front width 1.1, back width 1.3.

Legs. I-femur 12.0/ patella 5.0/ tibia 12.0/ metatarsus 11.0/ tarsus 6.5/ total 47.0; II- 11.0/ 5.0/ 11.0/ 10.0/ 6.0/ 43.0; III- 10.0/ 4.0/ 8.0/ 9.0/ 5.0/ 39.0; IV- 12.5/ 5.0/ 11.5/ 14.0/ 6.0/ 49.0. Cymbium 4.0 long, 2.6 wide, median apophysis 1.35 long.

Leg spination. Femora I p1-1-1-1, r1-1-1, d1-1- 1; II p1-1-1-1, r1-1-1-1-1, d1-1-1; III p1-1-1-1,r1- 1-1-1, d1-1; IV p1-1-1, r(2)-1-1-1, d1-1-1; patellae I-IV p1, r1; tibiae I-II v2 -2-2-2, p1-1, r1-1-1, d1-1-1; III-IV v2 -2-2, p1-1, r1-1, III d1-1, IV d1- 1-1-1; metatarsi I-II v2 -2-2, p1-1-1, r1-1-1; III v2 - 2-2, p1-2-1, r1-2-1; IV v2 -1-1-2-2, p1-2-1, r1-2-1; tarsi III-IV v2 -2-2-2-2. Palpal femora p1, r1, d1-1- 1-2; patellae p1; tibiae p1-1, r1.

Variation. Morphometric variability see Tab. 1. One male with a row of six spines dorsally on patella of one palp.

Female (SMNK 1447). Whole animal lightbrown, chelicerae dark brown to black, densely covered by long hairs, cheliceral boss reddishbrown. Abdomen with 2 pairs of dark spots on dorsum and 4 longitudinal lines of small dark spots on venter. Epigynum: median plate in its anterior part broad rectangular, prominent, with characteristic central triangular protuberance (Fig. 6), anterior border rounded or slightly invaginated (Fig. 29). Lateral plates broad, meeting the median plate near its anterior border. Epigynal openings situated anteriorly (Fig. 29). Copulatory ducts broad and straight, entering spermathecae from ventrally. Spermathecae nearly their diameter apart (Fig. 30).

Measurements. Total length 32.0. Prosoma 14.9 long, 11.6 wide, 3.5 high. Clypeus 0.9 high. Eye diameters and interdistances: AME 0.5, ALE 0.35, PME 0.6, PLE 0.6; AME-AME 0.5, AME-ALE 0.65, PME-PME 0.35, PME-PLE 0.8, AME-PLE 1.5, AME-PME 0.35, MOQ length 1.4, front width 1.15, back width 1.5. Chelicerae with 3 unequal promarginal and 4 subequal retromarginal teeth. Epigynal median plate 1.5 long, 1.05 wide.

Legs. I-femur 12.0/ patella 7.0/ tibia 9.5/ metatarsus 9.0/ tarsus 5.0/ total 42.5; II- 12.0/ 6.0/ 9.2/ 9.0/ 5.0/ 41.0; III- 10.5/ 5.0/ 9.0/ 9.0/ 5.0/ 38.5; IV- 12.0/ 5.0/ 12.0/ 12.5/ 6.0/ 48.0.

Leg spination. Femora I-II p1-1-1-1, r1-1-1, d1- 1-1; III p1-1-1-1, r1-1-1-1, d1-1-1; IV p1-1-1-1, r1, d1-2-2; patellae II p1; II -IV p1, r1; tibiae I-II v2 -2-2; III v2 -2-2, p1-1, r1-1, d1-1-1; IV v2 -2-2, p1-1-1, r1-1-1, d1-1-1; metatarsi I v2 -2-1, II v2 - 2-2, III v2 -2-1, p1-1-1, r1-1-1, d1-2-2; IV v2 -l-l- 2-2, p1-1-1, r1-1-1-1, d2-2-1-2; tarsi III-IV v2 -2- 2-2-2. Palpal femora p1, r1, d1-1-2; patellae p1; tibiae p1-1, d1; tarsi p2-1, r1.

Variation. See Tab. 2.

Coloration of living specimens (from São Paulo). Males lighter brown, cephalothorax with broad white marginal bands, and obscure diverging bands; abdomen dorsally with dark cardiac mark, 2 pairs of dark spots on anterior half and a pair of light spots on posterior half of abdomen. Metatarsi I and II dorsally whitish. Venter reddish brown with weak light stripes forming a triangle. Females generally darker brown. Cephalothorax without white bands, all other patterns less obvious than in male. All metatarsi dark.

Material examined. - BOLIVIA: Beni: San Ignacio. 5 [[males]] 1 [[female]] 2 imm., March 1961, L. Hammerschmidt, IBSP 1633. BRAZIL: Distrito Federal: Arredores de Brasília, 1 [[male]] 2 imm., Febr. 1971, W. Lourenço, IBSP 6501; Brasília, 1 [[male]], July 20 1995, Cadu, IBSP 8083; Brasília, 1 [[female]], May 5 1996, Cadu, IBSP 8087; Brasília, 1 [[female]], Dec. 1979, C. Carvalho, IBSP 4792; Brasília, 1 [[male]], April 13 1995, S.S. Sales, IBSP 6535; Lagoa Bonita, 1 [[female]], T.K. Brazil, UFBA, 1513; Poço Azul, 1 [[male]] 1 imm., April 1995, B. Duar et al, IBSP 6532; Poço Azul, 1 [[female]] 1 imm., April 1995, B. Duar et al, IBSP 6533; Poço Azul, 1 [[female]], Nov. 1994, R. Brandão, IBSP 6536; Goiás: Ipameri, 1 [[male]], May 1994, F.R. Alves, IBSP 5985; Jataí, Fazenda Aceiro, 2 [[males]] Oct. 1962, Exp. Depto. Zool., MZSP 11941; Leopoldina. 1 [[female]], July 1950, W.M. Silva, IBSP 430; Minaçu, Usina Hidrelétrica de Serra da Mesa, 1 [[male]], 1997, N.J. da Silva, IBSP 14642; Niquelândia, Serra da Mesa, 3 [[males]], Dec. 2 -22 1995, M. Rodrigues, IBSP 15012; Niquelândia, Serra da Mesa, 2 [[males]], Dec. 2 -22 1995, M. Rodrigues, IBSP 15013; Niquelândia, Serra da Mesa, 1 [[males]], Dec. 2 -22 1995, M. Rodrigues, IBSP 15014; Niquelândia, Serra da Mesa, 1 [[male]], Dec. 2 -22 1995, M. Rodrigues, IBSP 15020; Niquelândia, Serra da Mesa, Campinaçu, 1 [[female]], Febr. 18 - March 20 1996, M. Rodrigues, IBSP 15028; Niquelândia, Serra da Mesa Rio Bagagem, 5 [[males]] 3 [[females]], Sept. 24-30 1995, G. Skuk, IBSP 6272; Niquelândia, Serra da Mesa Rio Bagagem, 4 [[males]], Sept. 24-30 1995, G. Skuk, IBSP 6273. Tocantins: Ilha do Bananal, 1 [[male]], March 1961, V. Lenk, IBSP 5095; Palmas, 3 [[males]], March 11 -28 1998, M. Caleffo, IBSP 17577; Palmas, 2 [[males]], March 11 -28 1998, M. Caleffo, IBSP 17578. Minas Gerais: 2 [[females]], Aug. 17 1951, O. Figueiredo, IBSP 565; 1 [[female]], March 1980, P.A. Cabral, IBSP 2041; Barbacena, 1 [[female]], Nov. 1975, C. Haida, IBSP 4632; Belo Horizonte, 1 [[female]]. Oct. 1975, F.R. Segre, IBSP 4232; Belo Horizonte, Parque das Mangabeiras, 1 [[female]] and eggsac, Oct. 27 1983, G. Kistumacher, MNRJ 13036; Buritis, 1 [[female]], 1964, Exp. Depto. Zool., MZSP 4219; Cambuí, 1 [[male]], Nov. 12 1951, O. Figueiredo, MZSP 595; Fazenda Vale do Ipê, Campo Florida, 1 [[female]], Jan. 2 1968, J.G. Setto, IBSP 2096; Fronteira, 1 [[female]], Nov. 1982, J.H. Vieira, IBSP 3570; Heliodora, 1 [[female]], July 1986, A. Prat, IBSP 4995; Jacutinga, 1 [[male]], July 1982, P.A. Cabral, IBSP 3443; Lima Duarte, 1 [[male]], Oct. 1954, IBSP 1151; Mariana, 1 [[male]], May 1953, W. Wagner, IBSP 891; Ouro Preto, 1 [[female]], Febr. 1957, A. Nasser, IBSP 1380; Pirapetinga, 1 [[female]], Febr. 26 1995, R.S. Bérnilz, IBSP 6335; Rio Pandeiro, 2 imm., May 1935, J. Blaser, IBSP 982; Rio Vermelho, 1 [[female]], 15.11.95, Louzan & Vetanabaro, IBSP 20116; Santa Barbara do Mato Dentro, 1 [[female]], Oct. 1976, F.M. Arcano, IBSP 4784; Santa Rita do Jacutinga, 1 [[female]], Nov. 13 1936, O. Fonseca, IBSP 158. Pará: Caxiunã, Melgaço, 1 [[female]], Aug. 6 -16 1996, A.A. Lise, MCTP 8341; Serra do Cachimbo, 1 [[female]], Nov. 1955, Werner, MZSP 9708; Londrina, 1 [[female]], Oct. 1 1998, P.S. Bernadelle, MCTP 10359; Foz do Chopin, Cruzeiro do Iguaçu /Dois Vizinhos, 1 [[female]], Nov. 15 1998, Eq. IBSP, IBSP 21371; Campo Largo, 1 [[female]], March 19 1934, D.A. Mello, IBSP 147. Bahia: Itapé, 2 [[females]], Dec. 3 -17 1974, IBSP 161; Paulo Afonso, Hidrelétrica de Itaparica, 1 [[female]], Sept. 8 1988, T.K. Brazil, UFBA 234; Salvador, 1 [[male]], T.K. Brazil, UFBA, 1223. Mato Grosso: 1 [[female]], Sept. 1975, E. Romanini, IBSP 3647; Barra do Bugres, Estação Ecologica da Serra das Araras, 2 [[males]] 2 [[females]], Dec. 3 -9 1988, M.L. de B. Pereira et al., MNRJ 13043; 1 [[female]] 1 imm., Sept. 1993, C. Strussmann, MCTP 2994; Barra do Garças, 1 [[male]], Aug. 1976, A.T. da Costa, IBSP 6520; Cáceres, 2 [[males]], Jan. 1965, V. Veit, IBSP 1905; Chapada dos Guimarães, Buriti, 1 [[female]], Dec. 1992, L.F. Silva, MCTP 2954; 2 [[females]], MCTP 2956; Cuiabá, 1 [[male]], Oct. 11 1992, M. Detoni, MCTP 2998; Poconé 1 [[female]], Jan. 1982, R. Carsten, IBSP 3401; Rio Brilhante. 1 [[female]], June 18 1952, IBSP 722; Rio das Mortes, 1 [[male]] 2 [[females]] 1 imm., Aug.-Sept. 1949, Eq. IB, IBSP 574, 1244; Chupé, Rio das Mortes, 1 [[female]], 16.VII.1975, Equipe I.B., IBSP 3494; Santo Antonio de Leverger, 1 [[male]], Nov. 8 1982, M.I. Marques, MCTP 2543; Várzea Grande, 1 [[male]], June 1977, F.D. Hein, IBSP 2860; 1 [[female]], May 11 1989, IBSP 20120. Mato Grosso do Sul: Usina Hidrelétrica Sérgio Motta, Anaurilândia, 9 [[males]], 17 [[females]], Nov. 15 - Dec. 21 1998, Eq. IBSP, IBSP 21406; 1 [[female]], Sept. 1954, Cia. Mil. Manutenção, IBSP 953; 1 [[female]], Dec. 1981, C. Rollof, IBSP 3841; 1 [[female]], Jan. 1975, C.B.O. Santos, IBSP 4324; 1 [[male]], Jan. 1984, T. Fukasawa, IBSP 4678; 1 [[female]], Nov. 27 1975, IBSP 6517; Agachi, 5 [[males]] 4 [[females]], 1 imm., Aug. 14 - Oct. 31 1952, C.F. Moraes, IBSP 752, 760, 763, 788, 798, 802, 805, 810, 829; Aquidauana, 1 [[male]], Febr. 1980, R. Tsukamoto, IBSP 3841; Campo Grande, 1 [[female]], Jan. 1950, P. Bruno, IBSP 1114; Guaicurus, 1 [[male]], Aug. 25 1952, IBSP 768; Nioaque, 1 [[male]], Sept. 1974, C.B.O. Santos, IBSP 4323; Paranaiba, 1 [[female]] 1 imm., June 7 1984, IBSP 6523; Paranaiba, 2 [[males]] 2 [[females]], IBSP 6524; Paranaiba, 1 [[male]], IBSP 6525; Paranaiba, 2 [[males]] 3 [[females]], IBSP 6526; Paranaiba, 3 [[males]] 4 [[females]], 1983, R.R. da Silva, IBSP 6527; Paranaiba, 3 [[males]] [[female]], IBSP 6528; Paranaiba, 1 [[female]], IBSP 6538; Paranaiba, 2 [[females]], IBSP 6539; Paranaiba, 2 [[males]], 1984, IBSP 6540; Paranaiba, 1 [[male]] 1 [[female]], IBSP 6541; Paranaiba, 1 [[female]], IBSP 6542; Paranaiba, 1 [[male]], IBSP 6543; Paranaiba, 2 [[males]] 1 [[female]], IBSP 6544; Paranaiba, 1 [[male]] 2 [[females]], IBSP 6545; Paranaiba, 1 [[male]], Jan. 19 1984, IBSP 6546; Paranaiba, 1 [[female]], IBSP 6547; Paranaiba, 1 [[male]], May 7 1984, IBSP 6548; Ponta Porã, 1 [[female]], Oct. 1977, IBSP 2888; Três Lagoas, Fazenda Canaã, 1 [[female]], Nov. 1966, F. Lane, MZSP 5611; Corumbá, Passo da Lontra, 1 [[female]], May 22 1993, J. Raizer IBSP 20113; 1 [[female]], Nov. 27 1993, IBSP 20114; 1 [[female]], June 20 1993, IBSP 20115; 2 [[females]] 4 imm., May 14 1993, J. Raizer & E.S. Guimaraes, IBSP 11712; Monte Libano, Campo Grande, 1 [[male]], Oct. 26 1989, L. Miyaguska & E.A. Okamura, IBSP 20117; Campo Grande, Jardim América, 1 [[female]], Jan. 10 1985, J.G. Sierra, IBSP 20118; Campo Grande, Jardim Rita Vieira, March 27 1989, A. Correa, IBSP 20119; Coxim, 1 [[male]], Dec. 17 1986, E.G. Souza, IBSP 8381. Rio de Janeiro: 1 [[female]], Sept. 1981, TM. de Almeida, IBSP 4790; 60 km oeste de Niterói, 1 [[female]], 1961-1963, W. Bücherl, SMF; Rio de Janeiro, Barra Mansa, Campus SOBEU, 1 [[female]], Jan. 3 1997, E.F. Ramos, IBSP 13303; Barra Mansa, Campus SOBEU, 1 [[female]], Febr. 17 1997, E.F. Ramos, IBSP 13307; Barra Mansa, Campus SOBEU. 1 [[female]], July 2 1997, E.F. Ramos, IBSP 13309; Barra do Piraí, 1 [[male]], July 1950, J. Nobele, IBSP 978; Nova Friburgo, 1 [[male]] 4 [[females]], Aug. 1980, P. Mix, IBSP 1749, 2136, 3212; Nova Friburgo, 1 [[female]] 1 imm., P. Mix, IBSP 6507; Porciúncula, 3 [[females]], March 14 1934, T Brandão, IBSP 156, 162; Santo Antonio de Pádua, 1 [[male]], Nov. 1954, J.D. Silva, IBSP 1155; São José do Penedo, 1 [[male]], Oct. 1975, A. Torres, IBSP 4633; Vassouras, 1 [[female]], Aug. 5 1974, M.V Ferraz, IBSP 3232. São Paulo: Araraquara, 1 [[female]], Jan. 9 1997, Controle Zoonoses, IBSP 7542; Indaiatuba, 1 [[male]], Jan. 6 1997, B. Grohmann, IBSP 7533; São Lourenço da Serra, 1 [[male]], Nov. 1993, L.P. de Almeida, IBSP 7385; Sao Paulo: 1 [[female]], Oct. 1996, E.C. Mossolin, IBSP 7414; 1 [[female]], Dec. 10 1948, A.H. Streuli, IBSP 124; 2 [[males]], 1933, IBSP 146; 1 [[female]], Oct. 30 1950, T. Jy, IBSP 403; 1 [[female]] 1 imm., May 14 1951, F. Marques. IBSP 507; 1 [[female]], Aug. 6 1951, A.A. Schnal, IBSP 560; 1 [[female]], Dec. 26 1994, P. Regis, IBSP 721; 1 [[female]], Jan. 11 1995, K. Araujo, IBSP 725; 1 [[female]], March 1949, E. Vasquez, IBSP 1039; 1 [[male]] 1 [[female]], June 1950, L.I. Hertl, IBSP 1083; 1 [[male]] 1 [[female]], April 1950, J. Vicente. IBSP 1123; 1 [[male]], Nov. 1954, G. Righi, IBSP 1154; 2 imm., Dec. 1955, IBSP 1207; 1 [[female]], Sept. 1957, A. Streuli, IBSP 1346; 1 [[male]], Aug. 1963, E. Fernandes, IBSP 1845; 1 [[male]], July 1980, J.A.F. Veloso, IBSP 2224; 1 [[female]], Sept 1977, M.V. Granado, IBSP 2876; 1 [[female]], March 1982, S.A. Almeida, IBSP 3153; 1 [[female]], Oct. 1982, H. Fontolan, IBSP 3544; 1 [[female]] 1 imm., Oct. 1982, L.D. Vizotto, IBSP 3558; 1 [[female]], Dec. 1982, R. D'Avila, IBSP 3587; 1 [[female]], Dec. 1982, L.E. Vilalta, IBSP 3605; 1 [[female]], May 1983, J.L.E. Brunn, IBSP 3707; 1 [[female]], Aug. 1983, S. Diaféria, IBSP 3734; 1 [[female]], Dec. 1982, N.S. Macedo, IBSP 3780; 1 [[female]], July 1990, J.T. de Godoy, IBSP 5045; 1 [[male]], Nov. 1974, S. Purg, IBSP 5163; 1 [[male]], April 29 1991, IBSP 5305; 1 [[female]], March 1992, R.A. Rocha, IBSP 5797; 1 [[female]], Oct. 1990, Eq. IB, IBSP 5811; 1 [[female]], April 1969, K. Feher, IBSP 6004; 1 [[female]], Nov. 1993, L. P. de Almeida, IBSP 72412; 5 [[female]], April 30 1951, J. Eleutério, IBSP 492, 493; 4 [[females]] 2 imm., May 14 1951, F. Marques, IBSP 503, 505, 506; 1 [[female]], Oct. 26 1951, MZSP 11994; São Paulo, 1 [[male]], Oct. 26 1998, A.M. de Freitas, IBSP 20137; São Paulo, Horto Oswaldo Cruz, 1 [[female]], Aug. 4 1934, IBSP 145; Guarulhos, 1 [[female]], L.C. Strini, IBSP 729; Instituto Butantan, 1 [[female]], June 12 1962, IBSP 1769; Caxingui, 1 [[male]], Dec. 1962, I. Christe, IBSP 1816; Alto da Boa Vista, 1 [[male]], May 1978, K.P. von Robert, IBSP 6521; Campus Inst. Butantan, 1 [[female]], R.S. da Silva, IBSP 6502; Candido Rodrigues, 1 [[female]], Febr. 9 1952, S. Borghi, IBSP 641; Carlos Magalhães, 1 [[female]], Nov. 14 1951, A. Paulucci, IBSP 606; 1 [[female]], July 1995, IBSP 76925; São Paulo, 1 [[female]], MZSP 9225; Atibaia, 1 [[male]] 1 [[female]], April 1975, J.G. Oliveira, IBSP 2747; Avanhandava, 1 [[female]], 1990, E. Garbe, MZSP 5752; Avaré 1 [[female]], H. Gabriel, IBSP 6514; Bariri, 1 [[female]], Oct. 22 1951, R. Paulino, IBSP 592; Barretos, 1 [[female]], March 21 1952, IBSP 689; Bebedouro, 1 [[male]], Sept. 1957, A. Marques, IBSP 1345; Bragança Paulista, 1 [[male]] 2 [[females]], Oct. 1977, J.C. Korte, IBSP 2891; Bragança Paulista, 1 [[female]], Oct. 1981, I. Rodrigues, IBSP 6519; Cachoeira Paulista, 1 [[female]], Febr. 1980, L.A. Mello, IBSP 4625; Cachoeira do Marimbondo, 2 [[females]] 1 imm., 1964, Vizotto, MZSP 3384, 4461; Caieiras, 2 [[females]], May 1950, A. Streuli, IBSP 1121; Campinas, 1 [[female]], Aug. 1983, A. Schmidt, IBSP 3737; Campo Largo, Jarinú, 1 [[female]], Jan. 1980, A.C. Lopes, IBSP 6504; Catanduva, 1 [[female]], Nov. 6 1950, A. Magnollo, IBSP 409; Catiguá, 1 [[female]], Febr. 28 1949, IBSP 166; Cotia, 1 [[female]], June 1983, J.R.C. Pereira, IBSP 3723; Cruzeiro, 2 [[females]], Nov. 1973 - July 1974, L.C.A. Botelho, IBSP 2717; Santana de Parnaiba, Cururuquará, 1 [[female]], Jan. 1980, J. Rongett, IBSP 6515; Embuguassú, 1 [[male]], May 1979, K.P. von Robert, IBSP 6505; Embuguassú, 1 [[female]], April 1981, J.C. Pereira, IBSP 6516; Cipó, Embuguassú, 1 [[male]] 1 [[female]], May 1977, G. Longo, IBSP 6505; Embú, 2 imm., Jan. 1962, J. Avila, IBSP 1730; Embú, 1 [[female]], Nov. 1977, W. Silva, IBSP 6509; Engenheiro Margilac, 1 [[female]], Nov. 24 1977, N. Bianconi, IBSP 6508; Franco da Rocha, 1 [[female]], Nov. 6 1950, E. Rolland, IBSP 407; Gruta Mirassol, 1 [[female]], July 28 1966, Reis e Costa, MZSP 5124; Ilha Marimbondo, 2 [[females]], May 1975, H. Belluomiro, IBSP 3425; Ilha São Sebastião, 1 [[ male]], June 1950, H. Urban, IBSP 1143; Itapira, 1 [[female]], July 1982, L.A. Fernandes, IBSP 3407; Itú, 1 [[male]] 3 [[females]], April 1992, R. D'Avila, IBSP 2540, 5817; Itú, 1 [[female]], April 29 1997, R. D'Avila, IBSP 11713; Jarinu, 1 [[male]], March 1980, A.C. Lopes, IBSP 1573; Jundiaí, 1 [[female]], Nov. 25 1951, A. Pereira, IBSP 617; Juquitiba, 1 [[female]], Nov. 18 1995, A.C. Raci, IBSP 6393; Miracatu, 1 [[female]], S.A. Curtil, IBSP 731; Oriente, 1 [[female]], Oct. 15 1973, R. Giorgi, IBSP 2678; Osasco, 1 [[male]], March 29 1979, O. Calazans, IBSP 2915; Osasco, 1 [[female]], Jan. 16 1996, V.B. de Oliveira, IBSP 6537; Paraibuna, 1 [[female]], April 1983, M.A. Pochon, IBSP 3671; Parelheiros, 3 [[males]], April 1974, R. Schwarz, IBSP 2736; Pederneiras, 5 [[females]] 2 imm., Dec. 1958, F. Pinheiro, IBSP 1356; Pereira Barreto, Usina Hidreletrica Tres Irmãos, 1 [[male]], 7 [[females]] 12 imm., Sept.- Oct. 1990, Bertani, Costa & Bentin, IBSP 4865; 1 [[female]], Sept. 1990, SMNK 1447; 1 [[male]], Dec. 1 1986, SMNK 1448; 1 [[female]], SMNK 1449; 1 [[male]], SMNK 1450; Pindamonhangaba, 1 imm., Oct. 23 1950, L. Hertl, IBSP 376; Pindorama, 1 [[male]] 4 imm., Aug. 1950, A. Sartori, IBSP 1091; Pirassununga, 1 [[male]] 2 [[females]], Oct. 31 1940, Horaes e Machado, MZSP 6349; Poá, 1 [[female]], Oct. 1982, R. Tarteli, IBSP 3518; Ribeirão Preto, 1 [[male]], C. Neto, IBSP 3236; Rio Claro, Fazenda Paraguassú, Santa Gertrudes, 2 [[females]], June 2 1978, J.H.B. Medeiros, IBSP 6500; Rio Claro, 3 [[females]], Jan. 1943, F.S. Pereira, MZSP 6558; Roseira, 1 [[female]], R.V. França, IBSP 3666; Salto Grande, 1 [[male]], Aug. 27 1937, G. Calegari, IBSP 150; Santo Amaro, 1 [[male]], M.P. de Oliveira, IBSP 5978; São Paulo, 1 [[female]], M. de Souza, IBSP 6510; Serra dos Cristãos, Franco da Rocha, 1 [[female]], J.G.P. de Godoy, IBSP 6503; Socorro, 1 [[female]], IBSP 732; Suarão, 1 [[female]], Febr. 1983, O. Grubl, IBSP 3629; São José do Rio Preto, 1 imm., 1964, Vizotto, MZSP 4441; São Lourenço da Serra, 1 [[female]], L.P. de Almeida, IBSP 730; São Lourenço de Serra, 1 [[male]], Nov. 4 1993, L.P. de Almeida, IBSP 6408; São Luis do Paraitinga, 1 [[male]] 3 [[females]] 2 imm., Nov. 1983, L.C. Darte, IBSP 4676; São Miguel Paulista, 3 [[males]] 1 [[female]], March 1954, A. Castelicci, IBSP 925; São Roque, Fazenda São Joaquim, 1 [[female]], Jan. 1963, IBSP 1821; Taboão da Serra, 1 [[female]], Oct. 1982, R.A. Mateus, IBSP 3522; Tanquinho, 1 [[female]], Sept. 12 1934, L. Constancio, IBSP 148; Tatui, 1 [[female]], Oct. 1982, J.R. Pires, IBSP 3518; Tatui, 1 [[female]], July 19 1995, O.M. Martins, IBSP 6395; Tremembé 1 [[female]], July 1950, T. Miller, IBSP 1116; Usina Hidrelétrica de Agua Vermelha, Guarani, 2 [[females]], July 20 1978, G. Puorto, IBSP 6534; Usina Hidrelétrica de Rosana, 3 [[males]] 28 [[females]] 26 imm., Dec. 1986, Eq. IB, IBSP 4516; Vargem Grande Paulista, 1 [[male]], Nov. 4 1997, J. Bocco, IBSP 14483; near Rio Mogi Guaçu, 1 [[female]], June 1 1982, IBSP 5304; road to Parelheiros, Sitio Palmeiral, 2 [[males]] 2 [[females]], 1973, R. Schwarz, IBSP 2713; without locality, 1 [[male]] 2 [[females]], April 1956, A. Castelucci, IBSP 1280; Barueri, 1 [[female]], Dec. 15 1998, R.M. Fiorone, IBSP 20365; Barueri, 1 [[female]], July 1950, T Miller, IBSP 3445; Barueri, 1 [[female]], June 24 1961, K. Lenko, MZSP 7048; Presidente Epitácio, 5 [[males]] 1 [[female]] 2 imm., Jan. 1999, Eq. IBSP, IBSP 21405. Paraná: Foz do Iguaçu, Usina Hidrelétrica de Itaipú, 2 [[females]], Oct. 1982, Eq. IB, IBSP 1071; Porto Rico, 1 [[female]], May 14 1970, J.C. Corte, IBSP 8424; Ribeirão Claro, 1 [[female]], Jan. 1990, IBSP 5306. Santa Catarina: Barra do Sul, Balneário Barra do Sul, 1 [[female]], April 14 1996, E.M. Azeppa, IBSP 9528; Florianópolis, 1 [[male]] 2 [[females]], May 13 1997, M. Gerardi, IBSP 9995; Nova Teutonia, 2 [[females]] 1 imm., SMF 13869. Rio Grande do Sul: Pirapó, Rio Ijui-Mirím, 1 [[male]], Jan. 3 -12 1989, Eq. Garabi, MCTP 978; 1 [[female]], same data, MCTP 979; São Borja, Rio Uruguai, Jan. 3 -12 1989, Eq. Garabi, MCTP 981; Santo Angelo, 2 [[females]], Nov. 28 1951, O.L. Souza, IBSP 621. ARGENTINA: Without locality, 1 [[female]], Jan. 1971, A. Martinez, MACN 6464; Playa Nunez, 2 [[females]] 1 imm., Aug. 1936, D'Castillo, MACN. Buenos Aires: Delta del Parana, 1 [[male]] 1 [[female]], Febr. 1 1970, L. Yivo, MACN 6440. Chaco: Puerto Inpenetrable, 1 [[female]], Febr. 25 1973, Aguilera & Dominguez, MACN 6570; Resistencia, 1 [[female]], Dec. 1971, J. Aguilera, MACN 6419. Corrientes: Bela Vista, 1 [[female]], June 1962, Massoia, MACN 6158; Colonia, 1 [[male]], Febr. 1948, C. Pellegrini, MACN 6166. Delta: Rio Ibicuicito, 1 [[male]], Feb 1951, Tzabo, MACN 6171; Rio Ibicuicito, 1 [[female]], Dec. 1971, Tzabo, MACN 6172; Rio Carachay, 1 [[female]], July 1941, F. Monrás, MACN 6161. Formosa: Clorinda, 1 [[male]] 1 imm., Jan. 1949, Duret, MACN 4117; Guay, 1 [[female]], Sept. 1948, A. Pilaos, MACN 6632; Jujuy: Fraile Pintado, 1 imm., Jan. 1967, E. Maury, MACN 6035; Yuto, 1 [[female]], March 1967, Barrio, MACN 6162; Reserva Ecológica El Bagual, 26°10'53"S 58°56'39"W, Laishi, Dec. 1992 - March 1993, J.A. Corronca & C. S. Abdala (FMLT); 1 [[male]] 1 [[female]], same data (FMLT). Misiones: 30 km from Puerto Bemberg, 1 [[female]], Jan.-Febr. 1950, W. Partridge, MACN 3248; Candelaria, 1 [[female]], Dec. 1943, M.J. Viana, MACN 6157; Posada, 2 [[female]] 1 imm., Nov. 1962, Salguero, MACN 6177; Puerto Iguazu, 1 [[female]], Oct. 1954, De Carlo, MACN 6178; Parana de las Palmas, Isla del Rio Luján, 1 [[female]], July 1952, J. M. Gallard, MACN 3723. Salta: Pichanal, 1 [[female]], Dec. 1994, Schiapelli & Gerschman de Pikelin, MACN 6167; Urundel, 1 [[female]] 4 imm., Oct. 1948, Bachmann & Nunez, MACN 6181. Santa Fé: 1 [[male]], July 1963, Inali, MACN 6635; Las Gamas 20 km W of Vera, 2 [[females]] (eggsac), Oct. 30 1994, M. Ramirez & J. Faibovitch, MACN. Tucumán: 2 [[females]], Sept. 1948, M. Biraben, MACN 6168. PARAGUAY: Aregua, 1 [[female]] 1 imm., July 10 1980, NMNH; Aregua, 1 [[female]], June 28 1903, ZMB; Puerto Casado, 1 [[male]] 1 [[female]], Nov. 1994, Gia & Cramwell, MACN, 1593; San Bernardino: 1 [[male]], K. Fiebrig col., ZMB, 11433; 1 [[male]] 2 [[females]], K. Fiebrig col., ZMB, 1249/06; 3 [[females]], K. Fiebrig col., ZMB, 373/06.

Distribution. - Southern to central South America (Fig. 60)