Cephennium asturicum REITTER, 1880

Assing, Volker, 2021, On the Cephennium fauna of the Iberian Peninsula and the Atlantic Islands (Coleoptera, Staphylinidae, Scydmaeninae), Linzer biologische Beiträge 52 (2), pp. 891-931 : 916

Cephennium asturicum REITTER, 1880 ( Figs 7 , 31-33 , Map 3 )

Material examined: SPAIN: País Vasco: 1♂, Sierra de Cantabria, ca. 30 km S Vitoria, Puerto de Herrera, 42°36'19''N, 02°40'24''W, 1000 m, beech forest, litter and moss between rocks sifted, 14.X.2003, leg. Assing (cAss). La Rioja: 4♀♀, Sierra de la Demanda, S El Rio, 2.VIII.1971 (cAss). Cantabria: 18 exs., Picos de Europa, Valle de Salvoron , Espinama , 27.VII.1972, leg. Meybohm (cAss) ; 3 exs., same data, but 30.VII.1972 (cAss) ; 5 exs., Picos de Europa, Espinama to Puerto de Aliva , 29.VII.1972, leg. Meybohm (cAss) ; 20 exs., Santander, Ruente env., Monte Aa , 1.VIII.1972, leg. Meybohm (cAss) ; 1 ex., Reinosa env., Tres Mares-Cornon, 2000-2150 m, 4.VI.1991, leg. Wunderle (cAss). Asturias: 3 exs., Oviedo , Playa de la Franca , 31.VII.1972, leg. Meybohm (cAss) ; 2 exs., Picos de Europa, Covadonga, Refugio Vegarredonda , 1650 m, 7.VI.1991, leg. Wunderle (cAss) ; 3 exs., Picos de Europa, Lago de la Ercina , Vega Bricial , 43°15'N, 4°58'W, 5.VI.1998, leg. Lompe (cAss) ; 12 exs., Sierra de Rañadoiro, Puerto del Connio, 43°02'33''N, 6°43'08''W, 1300 m, N-slope, mixed forest on scree with birch, oak, Sorbus , and with Luzula and Erica undergrowth, 25.VI.2002, leg. Assing (cAss). Castilla y León: 1 ex., Picos de Europa , 40 km WNW Aguilar de Campoo, pass W Cervera, 42°53'52''N, 04°38'54''W, 1550 m, N-slope with beech and Sarothamnus, 13.VII.2003, leg. Assing (cAss) ; 7 exs., Caboalles, leg. Paganetti (cAss). Galicia: 6exs., Sierra de Ancares, ENE Degrada , 42°50'12''N, 6°54'07''W, 970 m, mixed deciduous forest with very old Castanea sativa, 14.VII.2004, leg. Assing (cAss) ; 1 ex., Sierra de Ancares, ESE Degrada, 42°48'53''N, 6°53'38''W, 1300 m, moist Ilex forest with ferns, 10.VII.2004, leg. Assing (cAss) .

Material examined by C. Besuchet (unpubl.): SPAIN: Navarra: 5 exs., Alsasua, leg. Daniel ( ZSM). Cantabria: 2 exs., Reinosa ( BMNH) ; 41 exs., Santander, Ruente env. , Monte Aa , leg. Franz ( MHNG, NHMW) ; 7 exs., Espinama env ., Coriscao, leg. Franz ( MHNG, NHMW) ; 11 exs., Espinama env., leg. Franz ( NHMW) ; 3 exs., Santander, Mte Saja near Saja, leg. Franz ( NHMW) ; 3 exs., Santander, Puerto del Escudo , leg. Franz ( NHMW). Asturias: 21 exs., Bezanes env. , Mte. Reres [?], 800-900 m, leg. Franz ( MHNG, NHMW) ; 11 exs., Cangas del Narcea , "Bosque d. Munielles [?]", leg. Franz ( NHMW) ; 2 exs., Picos de Europa , Peña Santa env., leg. Franz ( NHMW) ; 4 exs., Monte Montera, 800 m, leg. Franz ( NHMW) ; 5 exs., Covadonga, leg. Franz ( MHNG, NHMW) ; 2 exs., Puerto de Pajares , leg. Franz, Kricheldorff ( HNHM, NHMW) ; 9 exs. [two labelled as lecto- and paralectotype], locality not specified, leg. Getschmann ( HNHM, MCSNG, MNHNP, NHMW) ; 1 ex., locality not specified (coll. Stockmann).. Castilla y León: 113 exs., Caboalles, leg. Paganetti, Schatzmayr ( FMNH, HNHM, MHNG, MNB, NHMW, TLMF, coll. Mateu, coll. Perrot, and other collections) ; 3 exs., Burgos, Puerte de Carrales , leg. Franz ( NHMW, cAss) ; 5 exs., Ponferrada, leg. Paganetti ( MHNG, MNB). Galicia: 21 exs., Lugo, Valle de Lozera , leg. Franz ( MHNG, NHMW) ; 13 exs., Lugo, Sierra de Ancares , leg. Franz ( MHNG, NHMW). Locality ambiguous 2 exs., Peña Labra, leg. Daniel ( ZSM) .

Diagnosis: Body length 1.1-1.3 mm. Body broad and strongly convex in crosssection ( Fig. 7 ). Pronotum large in relation to elytra. Colouration usually reddish to reddish-brown.

Eyes composed of approximately 4-5 ommatidia. Pronotum approximately 1.25 times as broad as long and as broad as elytra; punctation extremely fine, barely noticeable. Elytra broad and relatively short; anterior impressions transverse and tomentose; supra-humeral carinae very fine; punctation more distinct than that of pronotum.

♂: protibia unmodified; mesotibia flattened and with dense long pubescence on inner face in apical third; metatibia weakly flattened on inner face in apical portion; aedeagus ( Figs 31-33 ) approximately 0.3 mm long; ventral process and internal structures of distinctive shapes; parameres relatively short, thin, straight, and with very long apical seta.

Distribution and natural history: This species is rather widespread in Northwest Spain, from Galicia in the west to Navarra and La Rioja in the east ( Map 3 ). The altitudes range from 800 to more than 2000 m.

Figures

Map 3: Distributions of Cephennium asturicum (black circles; based on Besuchet (unpubl.) and examined material) and of species of the subgenus Macroderus in the Iberian Peninsula: C. alacantae (black triangles); C. cujonicum (white diamond); C. segurae (black diamond); C. verrens (white circles); C. cazorlae (black square); C. nevadae (black star); C. malagae (white triangles).

Figs 7-11: Male habitus of Cephennium asturicum (7), C. filabresicum (8), C. espunae (9), C. tensum (10), and C. tenuissimum (11). Scale bar: 1.0 mm.

Figs 25-33: Cephennium kiesenwetteri (25-26), C. constrictum (27-30; 27-28: holotype; 29-30: paratype), and C. asturicum (31-33; 32-33: based on Besuchet unpubl.): (25-32) aedeagus in lateral, dorsal, and in ventral view; (33) internal structures of aedeagus in ventral view. Scale bars: 0.1 mm.