Porrhoclubiona Lohmander, 1944

Zhang, Jianshuang, Yu, Hao & Li, Shuqiang, 2021, On the clubionid spiders (Araneae, Clubionidae) from Xishuangbanna, China, with descriptions of two new genera and seven new species, ZooKeys 1062, pp. 73-122 : 73

Genus Porrhoclubiona Lohmander, 1944

Porrhoclubiona Lohmander, 1944: 20; Prószyński and Staręga 1971: 234; Sterghiu 1985: 54 (considered a subgenus); Wunderlich 2011: 140 (considered a genus); Marusik and Omelko 2018: 22 (elevated to genus).

Clubiona : Mikhailov 2012: 179 (synonymised Porrhoclubiona Lohmander, 1944).

Clubiona genevensis -group: Bosmans et al. 2017: 2.

Clubiona pteronetoides -group: Deeleman-Reinhold 2001: 96.

Type species.

Clubiona genevensis L. Koch, 1866 from Switzerland.

Diagnosis.

Distinct from all other clubionids by the cymbium with modified setae retrolaterally (not observed in P. pteronetoides Deeleman-Reinhold, 2001 Fig. 15B ); cymbial base with a tutaculum (Fig. 15B ); tegulum with a tegular groove serving as a conductor (Fig. 15D ); tibia with a strongly reduced prolateral apophysis (Fig. 15A ); subtegulum located posteriorly (Fig. 15C, D ); sperm duct U-shaped, with an additional loop located proximally to embolic base (Fig. 15D ); copulatory openings large, located posteriorly on ventral epigynal plate, united at midline, or separated but close (Fig. 16A, B ); epigynal plate with a somewhat protruded posterior margin (Fig. 16A, B ).

Description.

See Marusik and Omelko (2018) and Deeleman-Reinhold (2001).

Comments.

Porrhoclubiona is small Clubionidae with a relatively wide body and can be easily separated from Malamatidia , Matidia , and Nusatidia . Porrhoclubiona currently comprises two species groups, the Porrhoclubiona pteronetoides group and the Clubiona genevensis group. The Porrhoclubiona pteronetoides group was established by Deeleman-Reinhold (2001) based on two species endemic to the Oriental realm. The Clubiona genevensis group was formally named by Mikhailov (1992), though in fact it was first recognised by Lohmander (1944) as a subgenus of Microclubiona (currently considered junior synonyms of Clubiona ), and then refined by Bosmans et al. (2017) with eight species from West Palaearctic region. The two groups share almost all of the generic characters listed by Marusik and Omelko (2018) (see diagnosis above); however, Porrhoclubiona pteronetoides group can be differed from the Clubiona genevensis group by have a dorsal abdominal scutum (Fig. 16E, F ) (vs. absent), lack modified setae on the cymbium (Fig. 15B ) (vs. present) in males, and the spermathecae are elongate in females (Fig. 16C, D ) (vs. round).

Figures

Figure 15. Male palp of Porrhoclubiona pteronetoides A prolateral view B retrolateral view C bulb, prolateral view D bulb, ventral view E bulb, retrolateral view. Abbreviations: E = embolus; EB = embolic base; RTA = retrolateral tibial apophysis; TG = tegular groove; TU = tutaculum. Scale bars: 0.10 mm (equal for A, B, equal for C-E).

Figure 16. Porrhoclubiona pteronetoides, epigyne (A-D), male habitus (E, F) and female habitus (G, H) A intact, ventral view B cleared, ventral view C cleared, dorsal view D cleared, dorsal view E dorsal view F lateral view G dorsal view H ventral view. Abbreviations: BS = bursa; CD = copulatory duct; CO = copulatory opening; FD = fertilization duct; SP = spermatheca. Scale bars: 0.10 mm (equal for A-D); 1 mm (equal for E, F, equal for G, H).