Kamyristi exfrigore, Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024

Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent, 2024, Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities, Arthropod Systematics & amp; Phylogeny 82, pp. 151-181 : 151

3.2.1.1.1. † Kamyristi exfrigore sp. nov.

Figures 2G , 9A-C

Etymology.

" exfrigore " means "from the cold" in Latin as this species is the northernmost record of a megalyrid, extant and extinct, in a region where the temperature is currently below zero most of the year. The specific epithet is to be treated as an adjective.

Material studied.

Holotype female PIN 3730/411; housed in the collection of the Paleontological Institute of the Russian Academy of Sciences (PIN).

Type locality.

Baikura, Russia.

Age.

Upper Albian to lower Cenomanian, mid-Cretaceous.

Diagnosis.

Eye nearly round (vs. oval, higher than long in † Kamyristi yantardakhensis gen. et sp. nov.); axillae contiguous medially with axillar groove smooth (Fig. 9B ; vs. separated medially with axillar groove crenulated in † Kamyristi yantardakhensis gen. et sp. nov.).

Description.

Body length 2.03 mm. Almost complete, except ovipositor. - Head globular, higher than long (length 0.38 mm; height 0.43); slightly pubescent on vertex and frons; frons convex, shagreened; compound eye rounded, slightly higher than long, not covering head length, postocular carina absent; vertex convex, shagreened; torulus inserted very close to ventral margin of eye; subantennal groove present; mandible with three teeth; scape short and thick (length 0.09 mm); pedicel same length; flagellomeres cylindrical, longer than wide, elongate (length ca. 0.08 mm); flagellomere 12 longest; occipital carina crenulate. - Mesosoma shorter than metasoma (length 0.69 mm; height 0.40 mm), almost fully shagreened; mesoscutum shagreened, short (length 0.20 mm; width 0.47 mm), one third of mesosoma length, divided by crenulate median mesoscutal sulcus, with sparse erect setae; axillae continuous medially, with inner axillar groove smooth; mesoscutellum as long as mesoscutum, posterior margin slightly outcurved; pronotum not visible dorsally; anterior thoracic spiracle fully surrounded by pronotal cuticle; propodeum areolate-rugose. - Fore wing hyaline (length ca. 1.50 mm) and covered with microtrichia; C, Sc+R, Rs +M, basal segments of Rs and M; M+Cu, Cu and 1m-cu spectral; Rs not closing first submarginal cell and pigmented for two thirds between r-rs and Rs+M; M pigmented to level of 2r-m and then spectral to wing tip; pterostigma broad; R1 extending beyond marginal cell; Rs pigmented, arched toward stigma, closing marginal cell. - Legs bearing many scattered erect setae; one mesotibial spur, one metatibial spur present; metafemur swollen, metatibia notched before spur; metabasitarsus three times longer than following, remainder same length. - Metasoma almost half body length (length 0.96 mm), elongate, smooth; segments subequal in length; hypopygium well-developed, concave; ovipositor incomplete, covered by sheaths only at base; cerci triangular, exserted.

Figures

Figure 2. Drawings of fore wing venation for the taxa described in the present paper. A † Cretolyra noijebumensis gen. et sp. nov.; B † Cretolyra shawi gen. et sp. nov.; C † Genkyhag innebula gen. et sp. nov.; D † Megacoxa chandrahrasa gen. et sp. nov.; E † Megacoxa janzeni gen. et sp. nov.; F † Megacoxa synchrotron gen. et sp. nov.; G † Kamyristi exfrigore gen. et sp. nov.; H † Kamyristi yantardakhensis gen. et sp. nov. Scale bars: 0.5 mm (A - H).

Figure 9. † Kamyristi gen. nov. A - C † Kamyristi exfrigore gen. et sp. nov., holotype female PIN 3730 / 411, Albian-Cenomanian, Taimyr amber. A habitus in lateral view; B habitus in dorsal view; C head and mesosoma in lateral view; D - F † Kamyristi yantardakhensis gen. et sp. nov., holotype male 3311 / 2718, Santonian, Taimyr amber; D habitus in lateral view; E habitus in ventral view; E habitus in dorsal view. Scale bars: 0.5 mm (A, B, D, E); 0.25 mm (C). AG = inner axillar groove; ce = cercus; MMS = median mesoscutal sulcus; OC = occipital carina; OvS = ovipositor sheaths; pa = parameres; sp = anterior thoracic spiracle.