Tetragonoderus Dejean, 1829

Kavanaugh, David H., Cueva-Dabkoski, Mollie & Liang, Hongbin, 2023, Inventory of the Carabid Beetle Fauna of the Gaoligong Mountains, western Yunnan Province, China: Species of the Tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with Descriptions of Two New Species., Proceedings of the California Academy of Sciences 67 (20), pp. 451-491 : 463

Genus Tetragonoderus Dejean, 1829

Bembidium Gyllenhal, 1810:12 , in part [unjustified emendation for Bembidion Latreille, 1802 ; invalid name ( Bousquet 2012:535)].

Tetragonoderus Dejean, 1829:485 (type species Carabus quadrum Fabricius, 1792 , designated by Hope 1838:89); Bousquet 2012:1272.

Cyclicus Jeannel, 1949:870 (type species Tetragonoderus perrieri Fairmaire, 1900 , by original designation).

Metacyclicus Jeannel, 1949:871 (type species Tetragonoderus bastardi Alluaud, 1897 , by original designation).

Diagnosis. Adults of Tetragonoderus can be distinguished from those of Cyclosomus species in the study area by the following combination of character states: Body size smaller, BL = less than 6.0 mm; pronotum ( Figs. 13 B-F) narrower, widest at or anterior to middle; elytra slightly to distinctly and obliquely truncate apically, elytral color pattern varied; front tarsomeres 1 to 3 without distinct lateral expansions ( Figs. 14 B-F, 15B-F), broader in males than in females.

Diversity: At present, 16 species are recognized in the Asian fauna, and here we record four from the study area, including one new species.

Figures

FIGURE 13. Pronota of cyclosomines, dorsal aspect.A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (CASENT 1013512, Xiaojiangqiao, Qushi Township, Tengchong County, Yunnan, China); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov., (CASENT1039394, holotype, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT 1039412, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.5 mm.

FIGURE 14. Male front tarsus, dorsal aspect. A. Cyclosomus acutangulus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT 1012643, Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Tetragonoderus arcuatus Dejean (Nepalgunj, Banke District, Lumbini Province, Nepal); C. T. elegans Andrewes (CASENT1039499, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); D. T. parviculus Kavanaugh & Cueva-Dabkoski, sp. nov. (CASENT1039397, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); E. T. punctatus Wiedemann (CASENT1039548, Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China); F. T. microthorax Jian & Tian (8 km S of Jamiri, northeastern India). Scale lines = 0.2 mm.