Carlschoenherria gapudi Calcetas, 2019

Calcetas, Orlando A., 2019, Review of the genus Carlschoenherria (Coleoptera: Scarabaeidae: Melolonthinae) of the Philippines, Zootaxa 4619 (2), pp. 330-346 : 335-336

Carlschoenherria gapudi Calcetas new species

( Figs. 9–12 , 36 )

Type locality. Philippines, Mindanao Island .

Type material. Holotype (male), labeled: May 1993, V. Samarita ( NMM).

Description of holotype, male. Length 21.3 mm, width 8.7 mm. Body dichromatic, head, pronotum and scutel- lum blackish brown; anterior half of elytra dark reddish brown with blackish tinge, posterior half with dark aluta- ceous tinge, covered with yellowish white setae; leg dark alutaceous ( Fig. 9 ).

Head. Clypeus with anterior margin nearly straight dorsally.Antennomere III with short stalk, widely expanded apically; antennomere IV distinctly shorter than the rest of lamellae antennomeres. Lamellae length 2.5 mm, nearly as long as anterior margin of the clypeus. Terminal maxillary palpomere with distinctly long, shallow lanceolate shaped groove or depression, as long as antennomere VI. Mentum with indistinct latitudinal mesal carina, with shal- low depression below it; surface subrugosely punctate, covered with few setae, acicular-like scales.

Pronotum anterior margin concave, nearly straight mesally, weakly explanate, margin smooth mesally; margin towards anterior angle sloping upward, crenulate; anterior angle rounded, obtuse; margin crenulate, raised dorsally; lateral margin distinctly convex mesally, margin minutely crenulate, thinly carinate; posterior angle rounded, nearly at 90 0 angle, margin crenulate; posterior margin deeply concave mesally, smooth; surface rugosely punctate mesally and minutely punctate on each side; with narrow, shallow longitudinal groove or depression mesally, groove with long yellowish setae ( Fig. 10 ).

Scutellum anterior margin nearly straight; posterior margin widely concave or rounded apically; with few rounded punctures; surface with rounded, sparsely spaced punctures mesally, slightly rugged posteriorly.

Elytron with ten costae and ten intervals, each interval with dense, thick, stiff; setae longitudinally aligned along each interval; costae I–III wide, lustrous, impunctate; costae III–V coalesced apically; humeral margin distinctly wedge-shaped towards anterior angle; anterior angle rounded; lateral margin slightly convex mesally; apical margin widely rounded; sutural angle angulate, obtuse; sutural margin carinate near sutural angle and mesally, disappear- ing towards scutellum; surface covered with numerous, very narrow, elongate lanceolate to acicular-like, tapered apically white and yellowish setae.

Legs. Foretibiae with two teeth. Metatibiae with numerous long, short whitish scales; with numerous, sparsely distributed long brownish stiff setae; posterior metatibiae with 15–17 spicules.

Ventral side of thorax. Prosternal process spindle-shaped, slender longitudinally, truncate apically, tapered ba- sally. Metaventral process little prolonged, not reaching procoxae, attached mesally to the mesosternal process; metaventral process length 1.3 mm. Pro and mesosterna densely covered with long yellowish hairs except sparsely covered mesally.

Abdominal ventrites. Covered mostly with long hairs, long elongate lanceolate to acicular-like scales; abdomi- nal ventrites not uniformly covered with pubescence, lacking in pubescence at least near each anterior and posterior margins or rather narrowly restricted in the distribution of pubescence ( Itoh 1993); sides of abdominal ventrites II and IV with large oval shaped patches of scales or abdominal maculation; abdominal ventrite V with small patch of scales.

Pygidium. Anterior margin arched or convex mesally, posterior margin narrow.

Aedeagus. Posterior margin of parameres with elongate rounded protuberance, margin weakly concave towards lateral margin, longitudinally undulate towards apex. Length of apical process 1.0 mm from lateral view; apical margin bird-bill like laterally. Parameres and phallobase combined length 4.6 mm; as in Figs. 11, 12 .

Distribution. The Philippines (Mindanao Island).

Etymology. This new species is named after the late Dr. Victor P. Gapud, Professor Emeritus of Taxonomy and Systematics at the Crop Protection Cluster, University of the Philippines Los Baños. He is a versatile taxonomist working on different groups of Philippine insects such as Collembola, Odonata, Hemiptera and selected groups of Coleoptera .

Differential diagnosis. Based on the species-group classification of Itoh (1993), this species belongs to the C. sulcipennis species-group, the parameres are laterally compressed and bird-bill shaped at apex. Further, it can be classified into the C. philippinica species-subgroup, its pronotum has a longitudinal groove along the median line and the abdominal ventrites are lacking in setation at least near each anterior and posterior margins or rather narrowly restricted in the distribution of setation. In addition, the shape of the parameres resembles that of C. philippinica . In comparison to C. philippinica , antennomere III of C. gapudi Calcetas , new species is widely expanded apically and narrowly expanded in the former ( Fig. 10 ). Also, the setation and placement of patches of setae on the abdominal ventrites of C. philippinica and C. gapudi Calcetas , new species are similar ( Figs. 13, 14 ).

The collector failed to cite the locality of the specimen. However, examining the other melolonthine specimens collected by Dr. Samarita, he was collecting somewhere in the island of Mindanao during that time period.

Figures

FIGURE 36. Geographical distribution map of Carlschoenherria Bezděk, 2016 in the Philippines.

FIGURES 9–12. Carlschoenherria gapudi Calcetas, new species, holotype, male. 9, habitus, dorsal aspect; 10, antenna, lateral aspect; 11, genitalia, dorsal aspect; 12, genitalia, lateral aspect.

FIGURES 13–15. Abdomen, ventral aspect. 13, Carlschoenherria philippinica (Brenske), male; 14, Carlschoenherria gapudi Calcetas, new species, male; 15, Carlschoenherria sulcipennis (Laporte), male.