Vaccinium pseudobullatum W. P. Fang & Z. H. Pan

Nguyen, Thi Thanh Huong, Tagane, Shuichiro, Averyanov, Leonid V., Dang, Van Son, Fritsch, Peter W., Tong, Yi-Hua, Souladeth, Phetlasy & Nuraliev, Maxim S., 2025, Vaccinium bidoupense (Ericaceae), a new species from Vietnam and Laos, with notes on V. petelotii and V. pseudobullatum, PhytoKeys 266, pp. 75-96 : 75-96

Vaccinium pseudobullatum W. P. Fang & Z. H. Pan , Acta Phytotax. Sin. 19 (1): 108 (1981).

Fig. 2

Type.

China. Yunnan Province • Malipo County, Tiechang , 1200 m a. s. l., 20 February 1940, C. W. Wang 86988 [fl.] ( holotype: PE; isotypes: IBK, IBK 00302352 !; IBSC, 0457816 !; KUN, 12088909 ! & 12088910 !; SZ, 00126560 !; WUK, 0268189 , image!) .

Notes.

Based on the specimens cited in Appendix 1, we have clarified two androecial features of the species:

The length of the stamen filaments was indicated in the protologue ( Fang and Pan 1981) as 1–1.5 mm, which was repeated in Fang and Stevens (2005). Fang (1991) and Huang and Fang (1991) considered the range as 1–2 mm. In most of the specimens studied here, however, the filaments are 3–4 mm long. Both flowering gatherings listed by Fang and Pan (1981; Wang 86988 and Feng 13490) bear flower buds (and no fully open flowers), and we suppose it to be the reason that they indicate shorter filaments than those observed by us. Their description was likely reproduced in the subsequent account without additional investigation of flower structure.

The ratio of anther tubules to thecae was uniformly indicated as 2–3 for this species ( Fang 1991; Huang and Fang 1991, including fig. 105 (9, 10); Fang and Stevens 2005), whereas we observed it to be mostly 1.1–1.3. We therefore treat this feature as more variable than earlier thought.

Distribution.

S China (SE Yunnan, SW Guangxi), N Vietnam ( Cao Bang, Ha Giang). Until recently, the species was considered to be endemic to Yunnan; it was reported from Guangxi by Wei (2023) and from Vietnam by Averyanov et al. (2020).

Figures

Figure 2. Vaccinium pseudobullatum. A. Branch with leaves and flowers; B. Inflorescence, lateral view; C. Inflorescence, with front view of flowers; D. Bract and two bracteoles; E. Flower, lateral view; F. Androecium; G. Antepetalous (left) and antesepalous (right) stamens, dorsal view; H. Stamen, ventral view I. Three calyx lobes, ovary, disk and style; J. Cross section of ovary. Photos by Y. H. Tong, based on B. M. Wang TYH- 965 A.