Atholus torquatus ( Marseul, 1854 )

Cruz, Ian Niel dela, Hartini, Sri, Dwibadra, Dhian & Ôhara, Masahiro, 2024, Contribution to the Knowledge of the Genus Atholus (Coleoptera: Histeridae: Histerinae: Histerini) from the Indonesian Archipelago, Species Diversity 29 (1), pp. 119-139 : 136

Atholus torquatus ( Marseul, 1854)

( Figs 1G , 11C )

Hister torquatus Marseul, 1854: 587 (type locality: India).

Hister genuae Lewis, 1888: 639 [synonymized by Bickhardt (1913b: 698)].

Atholus torquatus : Lewis 1906: 402 (listed); Mazur 1984: 218; Mazur 1997: 134; Mazur 2011: 106 ( Nepal, Myanmar, Laos, Vietnam, Thailand, Indonesia: Java, Sumatra, China: Sichuan; catalogued); Mazur et al. 2015: 1454 ; dela Cruz and Ôhara 2023a: 8–16 ( Philippines).

Atholus genuae : Lewis 1906: 402 (listed).

Hister (Atholus) torquatus : Bickhardt 1910: 55 (catalogued); Bickhardt 1917: 194 (catalogued); Desbordes 1917: 325 (keyed).

Hister (Atholus) genuae : Desbordes 1917: 322 (keyed).

Specimens examined. OneJ, 4♀♀, 2 specimens of undetermined sex. Sumatra Island: West Sumatra, Bukit Gompong, Sukarami, NE 20 km from Padang , 19 August 1998, M . Ôhara leg . (1♀ SEHU); Central Sumatra, Payakumbuh, Harau Valley , November 1993, Sarimudanas leg . (1♀ MZB); Central Sumatra, Bukit Tinggi, Ngarai Sianok , 6 August 1997, T . Ito leg . (1J SEHU). Java Island: Baluran, Beokol and Banjuwangi , 16 May 1995, B . Pisarski and J . Proezyriski leg . (1♀ SEHU, 1 ex. MZB); Banten, Cidaon, Ujung Kulon , 06.4571°S, 105.1593°E, 11 March 2005, M . Ôhara leg . [JA-05-MO-013] (1♀ SEHU) . Borneo Island : West Kalimantan, Mt . Bawang, 250–300 m, March 1990, unknown collector [IC-22-in08] (1 ex. SEHU) .

Diagnosis. Atholus torquatus is distinguished by the combination of several characters: interrupted lateral pronotal stria, hamate on the apical end; absence of pronotal foveae near anterolateral angle and punctation on the basal half of pygidium, but becoming finer towards the apex.

Distribution. India, Nepal and China: Sichuan, Myanmar, Laos, Thailand, Vietnam, Philippines and Indonesia: Greater Sunda (Sumatra, Java and Borneo Islands) ( Fig. 11C ).

Remark. Two of the seven examined specimens exhibit the presence of the external subhumeral stria, an individual variability already pointed out by dela Cruz and Ôhara (2023a).

Figures

Fig. 1. Some Indonesian Atholus, dorsal habitus. A, Atholus bifrons (Marseul, 1854), IC-22-in01; B, A. philippinensis (Marseul, 1854), IC- 22-in06; C, A. coelestis (Marseul, 1857), IC-22-in02; D, A. myrmidon (Marseul, 1861), IC-22-in05; E, A. famulus (Lewis, 1892), IC-22-in03; F, A. singalanus (Marseul, 1880), IC-22-in07; G, A. torquatus (Marseul, 1854), IC-22-in08. Scale bar: 1.00mm.

Fig. 11. Map showing the occurrence of some species of Atholus from the Indonesian archipelago and its extra-limital distribution in the Oriental Region. A, Atholus coelestis; B, A. philippinesis; C, A. torquatus; D, A. bifrons.