Bakeriella Kieffer, 1910

Azevedo, Celso O., 2014, Synopsis of Bakeriella Kieffer, 1910 (Hymenoptera, Bethylidae), Zootaxa 3878 (6), pp. 501-535 : 502-504

publication ID	https://doi.org/ 10.11646/zootaxa.3878.6.1
publication LSID	lsid:zoobank.org:pub:F60A4F4D-49AF-4DAD-A60D-EB439A5A5DFD
DOI	https://doi.org/10.5281/zenodo.6126143
persistent identifier	https://treatment.plazi.org/id/60271C78-6514-4E16-FF15-FC680DBAFC8C
treatment provided by	Plazi
scientific name	Bakeriella Kieffer, 1910
status

Treatment

Bakeriella Kieffer, 1910 View in CoL

Bakeriella Kieffer 1910a: 288 View in CoL ; Kieffer 1910a (new species); Kieffer 1910b (new species); Evans 1964 (new species); Evans 1965 (new species); Evans 1966a (new species); Evans 1979 (key, new species); Evans & Fullerton 1979 (new species); De Santis 1981 (catalog); Gordh & Móczár 1990 (catalog); Azevedo 1991 (new species); Azevedo 1994 (key, new species); Kawada & Azevedo 2003 (new species); Azevedo et al. 2005 (new species).

Type species Bakeriella flavicornis Kieffer 1910a: 289 , by monotypy.

Diagnosis. Length varying from about 3.0 to 8.0 mm. Body predominantly black, few species with metallic reflection. Wings hyaline or subhyaline. Mandible with two or five apical teeth. Median clypeal lobe well projected forward. Lateral clypeal lobes usually inconspicuous. Median clypeal carina usually conspicuous and complete. Antenna with 11 flagellomeres. Occipital carina complete. Eye hairy. Notaulus present, usually almost complete posteriorly and widening posterad. Parapsidal furrow present, usually straight. Scutellar pits usually not connected by any groove. Metasoma well developed medially, scutellum and propodeal disc not in contact medially. Propodeal disc with three discal carinae, median carina complete, paramedian carina usually incomplete posteriorly, posterior corner usually conspicuously foveolate. Propodeal declivity with median carina. Forewing with three closed cells, (costal, median and submedian), radial vein always present and long. Mesopleuron with lower margin of lower fovea fused to episternal groove. Male genitalia: Paramere usually wide; digitus progressively narrowing apicad, apex acute, dorsal surface denticulate; aedeagus without constriction between apical and basal halves, ventral surface with conspicuous projection ventrad, apodeme usually thick ( Fig. 42 View FIGURES 36 – 50. 36 – 42 ).

Key to species

Females

1. Pronotal disc without carina ( Figs 34 View FIGURES 25 – 35. 25 – 28 , 51 View FIGURES 51 – 61. 51 – 55 ).................................................................. 2

- Pronotal disc with anterior carina ( Figs 25 View FIGURES 25 – 35. 25 – 28 , 89 View FIGURES 85 – 96. 85 – 88 ).............................................................. 6

2. Scutellar pits connected by wide groove; notaulus very short, absent on anterior half of mesoscutum ( Fig. 34 View FIGURES 25 – 35. 25 – 28 )............................................................................................... B. erythrogaster Evans

- Scutellar pits not connected by groove; notaulus present on anterior half of mesoscutum, at least partially ( Figs 48 View FIGURES 36 – 50. 36 – 42 , 70 View FIGURES 62 – 72. 62 – 64 )..... 3

3. Median clypeal lobe wide and rounded ( Fig. 121 View FIGURES 121 – 131. 121 – 123 ); head wider posteriorly ( Fig. 121 View FIGURES 121 – 131. 121 – 123 )................ B. quadriceps Evans

- Median clypeal lobe angulate ( Fig. 124 View FIGURES 121 – 131. 121 – 123 ); head wider medially ( Fig. 124 View FIGURES 121 – 131. 121 – 123 ).......................................... 4

4. Scutellar pit subrounded, only slightly wider than long ( Fig. 127 View FIGURES 121 – 131. 121 – 123 ).................................. B. reclusa (Evans)

- Scutellar pit transverse, distinctly wider than long ( Fig. 124 View FIGURES 121 – 131. 121 – 123 )................................................... 5

5. Subtegular mesopleural fovea opened ventrally; anterior mesopleural fovea closed ( Fig. 126 View FIGURES 121 – 131. 121 – 123 ).... B. quinquepartita (Kieffer)

- Subtegular mesopleural fovea closed; anterior mesopleural fovea opened posteriorly ( Fig. 91 View FIGURES 85 – 96. 85 – 88 )........... B. lindigi (Kieffer)

6. Pronotal disc without median carina ( Figs 48 View FIGURES 36 – 50. 36 – 42 , 148 View FIGURES 144 – 149. 144 – 147 )........................................................... 7

- Pronotal disc with median carina ( Figs 25 View FIGURES 25 – 35. 25 – 28 , 145 View FIGURES 144 – 149. 144 – 147 )............................................................. 14

7. Pronotal disc with lateral carina on anterior half ( Fig. 73 View FIGURES 73 – 84. 73 – 76 )...................................... B. inconspicua Evans

- Pronotal disc without lateral carina ( Fig. 56 View FIGURES 51 – 61. 51 – 55 )................................................................ 8

8. Scutellar pit rounded or subrounded ( Figs 56 View FIGURES 51 – 61. 51 – 55 , 148 View FIGURES 144 – 149. 144 – 147 )........................................................... 9

- Scutellar pit transverse ( Figs 14 View FIGURES 14 – 24. 14 – 16 , 48 View FIGURES 36 – 50. 36 – 42 )...................................................................... 11

9. Median clypeal lobe subtrapezoidal; eye very large ( Figs 6 View FIGURES 1 – 13. 1 – 5 , 148 View FIGURES 144 – 149. 144 – 147 ), VOL less than 0.5 × HE ( Figs 6 View FIGURES 1 – 13. 1 – 5 , 148 View FIGURES 144 – 149. 144 – 147 ); head with metallic reflection........................................................................................... 10

- Median clypeal lobe angulate ( Fig. 56 View FIGURES 51 – 61. 51 – 55 ); eye median-sized, VOL more than 0.5 × HE ( Fig. 56 View FIGURES 51 – 61. 51 – 55 ); head without metallic reflection................................................................................... B. grossensis Azevedo

10. Head rounded, frons without linear depression ( Fig. 6 View FIGURES 1 – 13. 1 – 5 ); scutellum without anterior groove ( Fig. 6 View FIGURES 1 – 13. 1 – 5 )................................................................................................... B. aurata Kawada & Azevedo

- Head elongate, frons with linear depression from clypeus to almost anterior ocellus ( Fig. 148 View FIGURES 144 – 149. 144 – 147 ); scutellum with inconspicuous groove connecting pits ( Fig. 148 View FIGURES 144 – 149. 144 – 147 ).............................................. B. sulcaticeps Kawada & Azevedo

11. Mandible with two apical teeth ( Fig. 15 View FIGURES 14 – 24. 14 – 16 ).................................................... B. brasiliana Evans

- Mandible with five apical teeth ( Fig. 49 View FIGURES 36 – 50. 36 – 42 ).................................................................. 12

12. Pronotal disc without posterior series of foveae ( Fig. 48 View FIGURES 36 – 50. 36 – 42 )........................................ B. floridana Evans

- Pronotal disc with posterior series of foveae ( Fig. 118 View FIGURES 109 – 120. 109 – 112 )....................................................... 13

13. Clypeal median lobe obtusely angulate ( Fig. 118 View FIGURES 109 – 120. 109 – 112 )................................................. B. polita Evans

- Clypeal median lobe narrowly truncate ( Fig. 107 View FIGURES 97 – 108. 97 – 101 )............................................... B. olmeca Evans

14. Median carina of pronotal disc very weak, almost imperceptible ( Fig. 141 View FIGURES 132 – 143. 132 – 135 )........................ B. subcarinata Evans

- Median carina of pronotal disc conspicuous ( Figs 70 View FIGURES 62 – 72. 62 – 64 , 82 View FIGURES 73 – 84. 73 – 76 )..................................................... 15

15. Temple strongly carinate ventrally ( Fig. 25 View FIGURES 25 – 35. 25 – 28 ).................................................. B. dentata Azevedo

- Temple not carinate ( Fig. 17 View FIGURES 14 – 24. 14 – 16 )........................................................................... 16

16. Carina of antennal scrobal carina almost longitudinal ( Fig. 17 View FIGURES 14 – 24. 14 – 16 )......................... B. bulbosa Azevedo & Moreira

- Carina of antennal scrobal carina mostly transverse ( Fig. 82 View FIGURES 73 – 84. 73 – 76 ).................................................. 17

17. Median carina of pronotal disc reaching anterior transverse carina ( Fig. 82 View FIGURES 73 – 84. 73 – 76 )................... B. lata Kawada & Azevedo

- Median carina of pronotal disc not reaching anterior transverse carina ( Figs 62, 70 View FIGURES 62 – 72. 62 – 64 )................................ 18

18. Median clypeal lobe narrowly truncate ( Fig. 70 View FIGURES 62 – 72. 62 – 64 )........................................... B. incompleta Azevedo

- Median clypeal lobe broadly rounded ( Fig. 62 View FIGURES 62 – 72. 62 – 64 ).................................................... B. inca Evans

Males

1. Pronotal disc ecarinate ( Figs 29 View FIGURES 25 – 35. 25 – 28 , 97 View FIGURES 97 – 108. 97 – 101 )....................................................................... 2

- Pronotal disc carinate ( Figs 1, 9 View FIGURES 1 – 13. 1 – 5 ).......................................................................... 5

2. Scutellar pits connected by wide groove ( Fig. 29 View FIGURES 25 – 35. 25 – 28 ).......................................... B. erythrogaster Evans

- Scutellar pits not connected by groove ( Figs 51 View FIGURES 51 – 61. 51 – 55 , 85 View FIGURES 85 – 96. 85 – 88 ).......................................................... 3

3. Paramere wide, less than 2 × as long as wide and paramere wider basally ( Fig. 101 View FIGURES 97 – 108. 97 – 101 )................ B. montivaga (Kieffer)

- Paramere slender, more than 2 × as long as wide and paramere wider apically ( Figs 55 View FIGURES 51 – 61. 51 – 55 , 88 View FIGURES 85 – 96. 85 – 88 )........................... 4

4. Ventral margin of paramere convex ( Fig. 55 View FIGURES 51 – 61. 51 – 55 )................................................... B. grandis Evans

- Ventral margin of paramere concave ( Fig. 88 View FIGURES 85 – 96. 85 – 88 )................................................. B. lindigi (Kieffer)

5. Pronotal disc without median carina ( Figs 92 View FIGURES 85 – 96. 85 – 88 , 135 View FIGURES 132 – 143. 132 – 135 )........................................................... 6

- Pronotal disc with median carina ( Figs 20 View FIGURES 14 – 24. 14 – 16 , 109 View FIGURES 109 – 120. 109 – 112 )............................................................. 15

6. Pronotal disc with lateral carina ( Figs 127 View FIGURES 121 – 131. 121 – 123 , 132 View FIGURES 132 – 143. 132 – 135 )............................................................. 7

- Pronotal disc without lateral carina ( Figs 9 View FIGURES 1 – 13. 1 – 5 , 43 View FIGURES 36 – 50. 36 – 42 )............................................................. 10

7. Scutellar pits rounded ( Figs 92 View FIGURES 85 – 96. 85 – 88 , 127 View FIGURES 121 – 131. 121 – 123 )...................................................................... 8

- Scutellar pits transverse ( Figs 73 View FIGURES 73 – 84. 73 – 76 , 132 View FIGURES 132 – 143. 132 – 135 )..................................................................... 9

8. Pronotal disc with foveolate thick groove U-shaped, lateral carina of pronotal disc very wide and incomplete on posterior third ( Fig. 92 View FIGURES 85 – 96. 85 – 88 ); paramere slender in lateral view ( Fig. 96 View FIGURES 85 – 96. 85 – 88 )............................................... B. mira Evans

- Pronotal disc without such groove, lateral carina of pronotal disc narrow and incomplete on posterior two thirds ( Fig. 127 View FIGURES 121 – 131. 121 – 123 ); paramere wide in lateral view ( Fig. 131 View FIGURES 121 – 131. 121 – 123 )..................................................... B. reclusa (Evans)

9. Head with vertex strongly convex ( Fig. 132 View FIGURES 132 – 143. 132 – 135 ); paramere evenly wide ( Fig. 135 View FIGURES 132 – 143. 132 – 135 ).......................... B. rossi Evans

- Head with vertex at least shortly straight medially ( Fig. 73 View FIGURES 73 – 84. 73 – 76 ); paramere slightly narrower basally ( Fig. 76 View FIGURES 73 – 84. 73 – 76 )..................................................................................................... B. inconspicua Evans

10. Scutellar pit rounded ( Fig. 1 View FIGURES 1 – 13. 1 – 5 ); paramere absent ( Fig. 4 View FIGURES 1 – 13. 1 – 5 )................................ B. absens Kawada & Azevedo

- Scutellar pit transverse ( Fig. 77 View FIGURES 73 – 84. 73 – 76 ); paramere present ( Fig. 81 View FIGURES 73 – 84. 73 – 76 )................................................... 11

11. Paramere very slender, at least 4.0 × as long as wide ( Fig. 117 View FIGURES 109 – 120. 109 – 112 )...................................... B. polita Evans

- Paramere wide, at most 2.0 × as long as wide............................................................... 12

12. Antennal scrobe not or weakly carinate ( Figs 43 View FIGURES 36 – 50. 36 – 42 , 65 View FIGURES 62 – 72. 62 – 64 ); paramere mostly evenly wide ( Figs 47 View FIGURES 36 – 50. 36 – 42 , 69 View FIGURES 62 – 72. 62 – 64 )..................... 13

- Antennal scrobe strongly carinate ( Figs 77 View FIGURES 73 – 84. 73 – 76 , 136 View FIGURES 132 – 143. 132 – 135 ); paramere with base distinctly narrower than median part ( Figs 81 View FIGURES 73 – 84. 73 – 76 , 139 View FIGURES 132 – 143. 132 – 135 ).. 14

13. Temple diverging anterad ( Fig. 43 View FIGURES 36 – 50. 36 – 42 ); antennal scrobe not carinate ( Fig. 43 View FIGURES 36 – 50. 36 – 42 ); paramere as long as basiparamere ( Fig. 47 View FIGURES 36 – 50. 36 – 42 )............................................................................................ B. floridana Evans

- Temple parallel ( Fig. 65 View FIGURES 62 – 72. 62 – 64 ); antennal scrobe weakly carinate ( Fig. 65 View FIGURES 62 – 72. 62 – 64 ); paramere longer than basiparamere ( Fig. 69 View FIGURES 62 – 72. 62 – 64 ).............................................................................................. B. incompleta Evans

14. Paramere sublozengy ( Fig. 140 View FIGURES 132 – 143. 132 – 135 ); aedeagus with apical half abruptly narrower ( Fig. 139 View FIGURES 132 – 143. 132 – 135 )............ B. rufocaudata Evans

- Paramere oval ( Fig. 81 View FIGURES 73 – 84. 73 – 76 ); aedeagus progressively narrowing apicad ( Fig. 80 View FIGURES 73 – 84. 73 – 76 )................ B. labans Azevedo & Moreira

15. Pronotal disc with lateral carina ( Figs 36 View FIGURES 36 – 50. 36 – 42 , 109 View FIGURES 109 – 120. 109 – 112 ); temple carinate ( Fig. 38 View FIGURES 36 – 50. 36 – 42 )......................................... 16

- Pronotal disc without lateral carina ( Figs 20 View FIGURES 14 – 24. 14 – 16 , 102 View FIGURES 97 – 108. 97 – 101 ); temple not carinate........................................... 17

16. Pronotal disc with anterior corner acute, median carina reaching its posterior margin ( Fig. 36 View FIGURES 36 – 50. 36 – 42 ); arms of cuspis almost equally wide ( Fig. 41 View FIGURES 36 – 50. 36 – 42 )........................................................................ B. flavicornis Kieffer

- Pronotal disc with anterior corner not acute, median carina not reaching its posterior margin ( Fig. 109 View FIGURES 109 – 120. 109 – 112 ); dorsal arm of cuspis much wider than anterior arm ( Fig. 112 View FIGURES 109 – 120. 109 – 112 )....................................................... B. olmeca Evans

17. Scutellar pits rounded ( Figs 20 View FIGURES 14 – 24. 14 – 16 , 144 View FIGURES 144 – 149. 144 – 147 )..................................................................... 17

- Scutellar pits transverse ( Figs 9 View FIGURES 1 – 13. 1 – 5 , 59 View FIGURES 51 – 61. 51 – 55 )...................................................................... 18

17. Propodeal disc with paramedian carina very wide and straight ( Fig. 20 View FIGURES 14 – 24. 14 – 16 ); mandible with two apical teeth ( Fig. 21 View FIGURES 14 – 24. 14 – 16 ); paramere slender and sinuous ( Figs 23, 24 View FIGURES 14 – 24. 14 – 16 )............................................................ B. cristata Evans

- Propodeal disc with paramedian carina narrow and arched ( Fig. 144 View FIGURES 144 – 149. 144 – 147 ); mandible with five apical teeth ( Fig. 145 View FIGURES 144 – 149. 144 – 147 ); paramere wide and not sinuous ( Fig. 147 View FIGURES 144 – 149. 144 – 147 ).......................................................... B. subcarinata Evans

18. Median carina of pronotal disc bifurcating anterad ( Fig. 9 View FIGURES 1 – 13. 1 – 5 )......................................... B. azteca Evans

- Median and anterior carinae with T-shaped junction, not bifurcating anterad ( Figs 59 View FIGURES 51 – 61. 51 – 55 , 102 View FIGURES 97 – 108. 97 – 101 ).......................... 19

19. Paramere progressively widened apicad ( Fig. 106 View FIGURES 97 – 108. 97 – 101 )..................................... B. nuda Kawada & Azevedo

- Paramere evenly wide ( Fig. 61 View FIGURES 51 – 61. 51 – 55 )................................................................ B. inca Evans