Skorikovia radoszkovskii ( Skorikov, 1935 )

Lelej, Arkady S., 2024, Revision of the Palaearctic genus Skorikovia Ovtchinnikov, 2002 (Hymenoptera: Mutillidae), Zootaxa 5529 (1), pp. 52-72 : 64-67

6. Skorikovia radoszkovskii ( Skorikov, 1935)

( Figs 19–27 , 39, 40 , 49 )

Mutilla quinquefasciata (non Olivier, 1811): Radoszkowski 1885: 15, fig. 11, ♂.

Ephutomma radoszkovskii Skorikov, 1935: 325 , ♂ ( Turkmenistan:Ashkhabad [Ashgabat], Repetek), lectotype, (designated Lelej et al. 2022: 73), ♂, “Caucasus [probably collected in Georgia] / Z / Type / 5-fasciata [handwritten] / Coll. Radosz[owski] / Zool. Muz. Berlin / Paralectotype Mutilla elongata Radoszkowski 1885 , B. Petersen des. 1988 / Smicromyrme radoszkovskii (Skorikov) , B. Petersen det. 1988 / Lectotypus Ephutomma radoszkovskii Skorikov Lelej design. 2011 / Skorikovia radoszkovskii (Skor.) Lelej det. 2011” [MNHU], examined.

Smicromyrme radoszkovskii : Lelej 1978: 81, ♂ [ Russia, North Caucasus: Stavropol Terr., Dagestan; Transcausus, North Iran].

Smicromyrme (Nemka) radoszkovskii : Lelej 1985: 244, ♂

Skorikovia radoszkovskii : Ovtchinnikov 2002: 95, ♂; Lelej 2002: 64, ♂; Ljubomirov 2006: 88, ♂ ( Turkey); Lelej & Yildirim 2009: 19 , ♂ ( Turkey); Lelej 2017: 155; Pagliano et al. 2020: 187, ♂.

Skorikovia pliginskiji : Yildirim 2006: 280, ♂ (Erzincan, Erzurum).

Diagnosis. MALE. Ocelli large, diameter of anterior ocellus almost 2.0 × distance between anterior and lateral ocellus. Posterior side of 2r-m cell thickened ( Fig. 26 ). Volsella with shallow basal emargination ( Figs 19 , 25 ). Apical third of hypopygium with median shallow emargination and lateral tuft of long setae. T1 length equal or less than its maximal width; dorso-basally without visible spiracle tubercles. Body length 8.0–12.0 mm. FEMALE. Pronotum medially black. Scutellar scale narrow, nail-like with prescutellar carina. T2 laterally with distinct pale setae. T4–5 with black setae. Pygidial area widened basally, coarsely sculptured with striae reaching posterior margin, apical part not glabrous. Body length 6.8–7.2 mm.

FEMALE (hitherto unknown). Body length 6.8–7.2 mm. Head. Ratio distance between eyes / long eye diameter 1.35 ×. Eyes elongated, slightly convex, not prominent in head profile. Clypeus basomedially with developed process. F1 equal to F2. Frons and vertex with yellowish setae not forming a distinct spot. Head behind eyes (dorsal view) slightly elongated. Head brownish red. Mesosoma . Mesosoma length 1.35 × mesonotum width; scutellar scale narrow, nail-like with prescutellar carina. Mesosoma and legs ferruginous, femora and tibiae brownish. Pronotum medially black. Mesosoma dorsally with sparse yellowish setae, black on anterior margin of pronotum. Metasoma. Metasomal segment 1 brown, posterior margin of T1 with fringe of pale setae. T2 basally with medial and distinct lateral spots of pale setae. apically with band of pale setae strongly widened medially. T3 with band of pale setae, T4–5 with black setae, T4 with lateral tuft of whitish setae. T6 with lateral tuft of brown setae. Pygidial area widened basally, rounded apically, coarsely sculptured with striae reaching posterior margin. Metasoma ventrally with sparse pale setae, S2–5 with fringe of pale setae.

Material examined. (45♂, 2♀). Type material. Lectotype of Ephutomma radoszkovskii Skorikov. Additional material. Turkey: Erzurum: 1850 m, 22.VII.1995, 1♂, E. Yildirim leg. [ EMET]; Dereboğazı , 2070 m, 11.VIII.1994, 1♂, E. Yildirim leg. [ IBSS]; Konaklý , 2000–2400 m, 22.VII.2000, 3♂, E. Yildirim leg. [ EMET]; Oltu , Vocational Training School, 1345 m, 20.VIII.2006, 1♂, E. Yildirim leg. [ IBSS]; Şenkaya , Turnalı, 1400 m, 8.VIII.1997, 2♂, E. Yildirim leg. [ EMET]. NevŞehir: Uçhisar , Kapadokia, 1450 m, 23–24.VII.1995, 2♂, J. Gelbrecht & E. Schwabe leg. [ IBSS]. Georgia: Vardzia , 1.IX.1976, 1♂, E. Narchuk leg. [ IBSS]; Azerbaijan: Adzhikend , 12.VII.1909, 1♂, Babadzhanidi [ ZIN]. Russia: Dagestan, Petrovsk-port [Makhachkala], 27.VII–1.VIII.1926, 4♂, M. Ryabov [ ZIN]; same place, 10, 17, 25.VII, 13.VIII.1926, 4♂, M. Ryabov [ ZIN]; Kizlyar , Yaman-aul, sandy area, 18.VII.1927, 1♂, A. Kirichenko [ ZIN]; Kumtorkale [near Makhachkala], 6.VII.1927, 1♂, M. Ryabov [ ZIN]; Aleksanrdo-Nevskoe , 21.VI.1927, 1♂, G. Olsoufieff [ ZIN]; Belidzhi Station , 15.IX.1926, 23.VII.1933, 4♂, M. Ryabov [ ZIN]; 27 km SSE Kochubei, on light, 21–22.VII.2015, 4♂, V. Loktionov, M. Proshchalykin, M. Mokrousov [ IBSS]; 4 km N Almalo, 43°08′19″N 47°12′42″E, 25.VI.2018, 14♂, Yu. Astafurova, et al. [ IBSS]; 10 km W Aglobi, Kamyshchai River valley , 41°54′29″N 48°13′59″E, 29.VI.2018, 1♀, Astafurova et al. [ IBSS]; Kamyshchai River valley , 41°54′29″N 48°13′59″E, 10.VI.2017, 1♀, M. Mokrousov [ IBSS] .

Distribution. Russia (Dagestan), Azerbaijan, Georgia, Turkey (Asian territory).

Remarks. For Mutilla quinquefasciata sensu Radoszkowski 1885 the new name Ephutomma radoszkovskii was proposed by A. Skorikov (1935) on the basis of the Radoszkowski genitalia figure ( Radoszkowski 1885: Fig. 11 ) with distribution in Turkmenistan (Ashgabat and Repetek) ( Skorikov 1935). In the Mutillidae collection of ZIN, studied by A. Skorikov, I could not find the specimens of Skorikovia with such genitalia from Ashgabat, where it can be mixed with S. elongata , and from Repetek, where Skorikovia species are not distributed. The specimens from the North Caucasus ( Lelej 1978) have the genitalia shown by Radoszkowski (1885) and I considered them to be true E. radoszkovskii . Furthermore, the lectotype [MNHU] has the handwritten label O. Radoszkowski “ 5-fasciata ”. With this lectotype designation I validated the use of Skorikovia radoszkovskii ( Skorikov, 1935) . The paralectotype label “ Mutilla elongata Radoszkowski 1885 , B. Petersen des. 1988” is not valid because O. Radoszkowski described and illustrated Mutilla quiquefasciata and M. elongata in one paper ( Radoszkowski 1885) and he could not use the same specimen for two species.

A long series of males of Skorikovia radoszkovskii has been collected from southern Dagestan. In this area the genus Skorikovia is represented by only one species. The female of S. radoszkovskii may eventually be recognized as the opposite sex of this species, based on their co-occurrence in southern Dagestan. There is no direct evidence (pair collected in copula) to support this relationship.

Figures

FIGURES 11–19. Skorikovia, males. 11–14. S. anatolica, holotype; 15. S. trinotata (paratype of pliginskiji), Crimea; 16, 17. S. elongata, Betpakdala, Central Kazakhstan; 18. S. transcaucasica, holotype; 19. S. radoszkovskii, Dagestan, Makhachkala. 11, 17. Forewing; 12, 15, 16. Genitalia (right part ventral view, left part dorsal view). 13. Head; 14. Scape, pedicel and F1–3. 18, 19. Volsella, dorsal view. (11–14 from Lelej & Yildirim 2009; 15–19 from Lelej 1985).

FIGURES 20–27. Skorikovia radoszkovskii, ♂. 20–24. Dagestan; 25. Georgia; 26, 27. Lectotype, Caucasus. 20. Habitus, dorsal view; 21. Habitus, lateral view; 22. Head, face view; 23. S8, ventral view; 24. Genitalia, dorsal view; 25. Genitalia, ventral view; 26. Forewing; 27. Labels.

FIGURES 35–43. Skorikovia, females. 35, 36, S. elongata, South-East Kazakhstan; 37, 38, S. maxim sp. nov., paratype; 39, 40, S. radosdzkovskii, Dagestan; 41, 42, S. trinotata, Volgograd, Russia; 43, S. serta, lectotype, montage. 35, 37, 39, 41, 43, dorsal view; 36, 38, 40, 42, lateral view.

FIGURES 44–55. 44, 47. Skorikovia elongata, female, South-East Kazakhstan; 45, 48. S. maxim sp. nov., female, paratype; 46, 50. S. trinotata, female, Volgograd; 49. S. radosdzkovskii, female, Dagestan; 51, 52. S. serta, female, lectotype. 53–55. S. sanguinicollis, male, holotype. 44–46. Head, face view; 47–51. T6, dorsal view; 52, 55. Labels. 53. Habitus, lateral view; 54. Genitalia, ventral view.