Manota Williston, 1896

Hippa, Heikki, 2006, Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species, Zootaxa 1161 (1), pp. 1-49 : 7-10

publication ID	https://doi.org/ 10.11646/zootaxa.1161.1.1
publication LSID	lsid:zoobank.org:pub:A1C3B9CA-8A9D-4B41-9F9D-81BFC31CFB2D
persistent identifier	https://treatment.plazi.org/id/5D4C8785-2842-FFA6-5C1A-0AA2FAEFF956
treatment provided by	Felipe
scientific name	Manota Williston
status

Treatment

Key to the Oriental and Palaearctic species of Manota Williston View in CoL View at ENA

This key is based only on males. For M. orientalis , the thoracic chaetotaxy is based on the holotype female, and the characters of the male hypopygium on the illustration by Edwards (1928).

1. Laterotergite setose........................................................................................................2

Laterotergite nonsetose .............................................................................................. 12

2. Preepisternum 2 (katepisternum) setose; tergite 9 distinct, short, extending posteriorly at most to half length of gonocoxae on ventral side; gonocoxa with position III megaseta/e present................................................................................................................... 3

Preepisternum 2 (katepisternum) nonsetose; tergite 9 indistinct, united laterally with gonocoxae, extending posteriorly as far as gonocoxae on ventral side; gonocoxa with position III megaseta/e absent....................................................................................... 8

3. Tegmen unusual, long, extending more posteriorly than gonocoxa, its median part strongly constricted giving tegmen a spoon or spadelike appearance ( Fig. 3B View FIGURE 3 ) .......... ............................................................................................................... M. spadix View in CoL sp. n.

Tegmen normal, short, extending less posteriorly than gonocoxa, elongatesubtriangu lar in shape (e.g., Fig. 3D View FIGURE 3 ) ............................................................................................ 4

4. Gonocoxa in position III with 1 stout megaseta; parastylar lobe absent ( Figs. 4B, D View FIGURE 4 ) 5

Gonocoxa in position III with 2 slender megasetae; parastylar lobe present ( Figs. 5B, E View FIGURE 5 ) ................................................................................................................................... 7

5. Rounded lobe on dorsal mesial margin of gonocoxa with few subequal normal setae ( Fig. 3D View FIGURE 3 )................................................................................................. M. roslii View in CoL sp. n.

Rounded lobe on dorsal mesial margin of gonocoxa with few subequal normal setae and long megaseta ( Figs. 4B, D View FIGURE 4 ) .................................................................................. 6

6. Stronger dorsallydirected setae subapically on ventral mesial margin of gonocoxa arising from small lobe, all setae in 1 group ( Fig. 4D View FIGURE 4 ) .......................... M. pollex View in CoL sp. n.

Stronger dorsallydirected setae subapically on ventral mesial margin of gonocoxa not arising from lobe as margin is evenly curved, 1 seta displaced from group more laterally ( Fig. 4 B View FIGURE 4 ) ........................................................................................... M. yongi View in CoL sp.n.

7. Gonostylus elongateoval, without mesial subapical lobe ( Fig. 5B View FIGURE 5 ) ....... M. ovata View in CoL sp.n.

Gonostylus narrow, almost parallelsided on basal threequarters, with mesial subapical lobe ( Fig. 5E View FIGURE 5 ) .............................................................................. M. angustata View in CoL sp. n.

8. Gonostylus bilobed; gonocoxa with 3 + 1 megasetae in position I ( Figs. 5B, C View FIGURE 5 ) .......... ................................................................................................................. M. biloba View in CoL sp.n.

Gonostylus onelobed (e.g. Fig. 6E View FIGURE 6 ); gonocoxa with 7 or more strengthened setae or megasetae in position I in straight or curved comblike row ( Figs. 6E View FIGURE 6 ; 7B, E View FIGURE 7 ; 8C View FIGURE 8 ) .... 9

9. Gonocoxa with 7 megasetae in position I; apex of megasetae roundly expanded ( Fig. 6F View FIGURE 6 ) ........................................................................................ M. heptacantha View in CoL sp. n.

Gonocoxa with 10 or more strengthened setae in position I; apex of setae sharp ( Figs. 7B, E View FIGURE 7 ; 8C View FIGURE 8 ) 10

10. Gonocoxa apicolaterally with exceedingly long and strong sigmoid setae ( Fig. 7E View FIGURE 7 ) .... ..................................................................................................................... M. ulu View in CoL sp. n.

Gonocoxa apicolaterally with short normal setae only ( Figs. 7B View FIGURE 7 , 8B View FIGURE 8 ) ...................... 11

11. Gonocoxa with 2 strong setae in position II, setae equal in size to those in position I, gonocoxal margin roundly angulate at this point ( Figs. 7B, C View FIGURE 7 ) .......... M. simplex View in CoL sp. n.

Gonocoxa without strong setae in position II, gonocoxal margin evenly curved at this point ( Fig. 8C View FIGURE 8 ) ...................................................................................... M. clausa View in CoL sp. n.

12. Katepisternum setose .................................................................................................. 13

Katepisternum nonsetose ........................................................................................... 32

13. Gonocoxa dorsally with long, apicolaterally attenuating and posteriorly directed setigerous lobe which is contiguous with lateral margin of gonocoxa, the lobe extending much more posteriorly than mesial or apicomesial margin of gonocoxa (e.g., Figs. 8E, F View FIGURE 8 ).....................................................................................................................................14

Gonocoxa dorsally without long apicolateral lobe (e.g., Figs. 14C, F View FIGURE 14 ), but if lobe is present in this position, it is transverse and with basal part wholly overlapped by apicomesial margin of gonocoxa (e.g., Fig. 15C View FIGURE 15 ) ........................................................... 24

14. Gonocoxal megasetae in position III directed obliquely anteriad ...... M. delyorum Papp View in CoL

Gonocoxal megasetae in position III directed transversely or obliquely posteriad (e.g., Fig. 8E View FIGURE 8 )........................................................................................................................ 15

15. Gonostylus with apicolateral angle and straight, obliquely truncated apex ( Figs. 8E View FIGURE 8 ; 9B, D View FIGURE 9 ; 10A, B, C View FIGURE 10 )........................................................................................................ 16

Gonostylus without apicolateral angle, entire apex rounded (e.g., Figs. 11B, E View FIGURE 11 ) ....... 18

16. Gonostylus short, mesial ventral margin convex, apicomesial angle produced as small lobe ( Figs. 8E View FIGURE 8 , 10B View FIGURE 10 ) ......................................................................... M. calcarata View in CoL sp. n.

Gonostylus long, mesial ventral margin concave, apicomesial angle not produced ( Figs. 9B, D View FIGURE 9 ; 10A, C View FIGURE 10 ) ................................................................................................. 17

17. Lateral margin of gonostylus nearly straight, apex of gonostylus much broader than medial width of gonostylus, setae at apicomesial angle of gonostylus in longitudinal rows ( Figs. 9B View FIGURE 9 , 10A View FIGURE 10 ) ........................................................................... M. procera View in CoL sp. n.

Lateral margin of gonostylus convex, parallel with concavity of the mesial margin, apex of gonostylus only slightly broader than medial width of gonostylus, setae at apicomesial angle of gonostylus arranged in 2 short transverse rows on dorsal side ( Figs. 9D View FIGURE 9 , 10C View FIGURE 10 ) ......................................................................................... M. transversa View in CoL sp. n.

18. Gonostylus with 1 or 2 setose mesial lobes ( Fig. 11E View FIGURE 11 , Edwards 1928: Fig. 2a View FIGURE 2 ) ........ 19

Gonostylus without mesial lobes (e.g., Fig.11B View FIGURE 11 ) ....................................................... 21

19. Gonostylus with 2 small mesial lobes, 1 on basal half, the other on apical half (E dwards 1928: Fig. 2a View FIGURE 2 ) .......................................................... M. orientalis SeniorWhite View in CoL

Gonostylus with 1 large mesial lobe ( Fig. 11E View FIGURE 11 ) ......................................................... 20

20. Dorsal mesial margin of gonocoxa acutely angulate subapically ( Fig. 11F View FIGURE 11 ) ................. ........................................................................................................ M. acutangula View in CoL sp. n.

Dorsal mesial margin of gonocoxa rounded subapically .................. M. bilobata Papp View in CoL

21. Gonostylus straight ( Figs. 11B View FIGURE 11 , 12D View FIGURE 12 ).......................................................................... 22

Gonostylus curved ( Figs. 12A View FIGURE 12 , 13D View FIGURE 13 ) ......................................................................... 23

22. Dorsal mesial margin of gonocoxa subapically with angulate lobe with row of marginal setae ( Figs. 11B, C View FIGURE 11 ); subbasal mesial setae of gonostylus all similar to other ventral and mesial setae of gonostylus ( Fig. 11B View FIGURE 11 ) ................................ M. pectinata View in CoL sp. n.

Dorsal mesial margin of gonocoxa subapically rounded ( Fig. 12 E View FIGURE 12 ); gonostylus with 2 subbasal mesial setae that are stronger than other ventral or mesial setae of gonostylus ( Fig. 12 D View FIGURE 12 ) ........................................................................................... M. ferrata View in CoL sp. n.

23. Gonostylus broad, 3 times as long as wide; sternite 10 large, halves with ca. 30 setae ( Fig. 12B View FIGURE 12 ) ................................................................................................. M. fera View in CoL sp. n.

Gonostylus narrow, over 5 times as long as wide; sternite 10 small, halves with ca. 15 setae ( Fig. 13D View FIGURE 13 ) .................................................................................... M. curvata View in CoL sp. n.

24. Gonocoxa dorsally in position I with group of blunt megasetae ( Fig. 13B View FIGURE 13 ) .................. .......................................................................................................... M. globigera View in CoL sp. n.

Gonocoxa with the usual sharp setae in position I (e.g., Figs. 14C, F View FIGURE 14 ) ...................... 25

25. Gonostylus with large, trapezoidal, nonsetose, striated plate on mesial side ............... ........................................................................................................ M. chinensis Šev View in CoL ćí k

Gonostylus without nonsetose striated mesial plate ................................................. 26

26. Dorsal mesial margin of gonocoxa produced as large lobe on apical part ( Fig. 14C View FIGURE 14 ) 27

Dorsal mesial margin of gonocoxa straight or evenly rounded (e.g. Fig. 15C View FIGURE 15 ) or with only small steplike discontinuity ( Fig. 15F View FIGURE 15 ) ............................................................. 28

27. Gonostylus 1.5 times as long as wide; parastylar lobes present; setae on position III arising from long common basal body ( Fig. 14B View FIGURE 14 ); lobe on dorsal mesial margin of gonocoxa narrow, without strong setae on posterior margin ( Fig. 14C View FIGURE 14 ) ........................ ............................................................................................................ M. cerciflex View in CoL sp. n.

Gonostylus nearly 3 times as long as wide; parastylar lobes absent; setae on position III arising nearly separately ( Figs. 14E, F View FIGURE 14 ); lobe on dorsal mesial margin of gonocoxa broad, with strong setae on posterior margin ( Fig. 14F View FIGURE 14 ) ......................... M. pappi View in CoL sp. n.

28. Gonostylus long and narrow, 4 or more times as long as broad, curved on apical part ( Figs. 15B, E View FIGURE 15 ) .............................................................................................................. 29

Gonostylus short and broad, at most 3 times as long as broad, not curved on apical part ( Figs. 16B, D View FIGURE 16 ; 17B View FIGURE 17 )..................................................................................................... 30

29. Gonostylus with long apical megaseta and with row of long flattened setae apicodorsally; setae in positions III and IV with short basal bodies ( Fig. 15B, C View FIGURE 15 ) ...................... .............................................................................................................. M. horrida View in CoL sp. n.

Gonostylus without apical megaseta, with normal setae apicodorsally; setae in positions III and IV with very long basal bodies ( Figs. 15E, F View FIGURE 15 ) ................ M. duplex View in CoL sp. n.

30. Gonostylus at basomesial corner with 1 or 2 strong and long setae deviating from other marginal setae; setae on position III arising from basal body that is several times longer than broad,and dorsal to this,in position IV,with similar body with1seta( Fig.16B View FIGURE 16 ) .......................................................................................................... M. perpusilla View in CoL sp. n.

Gonostylus at basomesial corner with setae similar to its other marginal setae; setae in position III with short basal body that is only twice as long as broad basally, dorsal to this, in position IV, with short oblique lobe with several setae ( Figs. 16D, E View FIGURE 16 ) .......... 31

31. Gonostylus twice as long as broad, with small mesial lobe just in apical half with long straight seta,the mesial margin of gonostylus with ca. 5long setae on basal half ( Fig. 16D View FIGURE 16 ) ....................................................................................................... M. oligochaeta View in CoL sp. n.

Gonostylus ca. 2.5 times as long as broad, without lobe on mesial margin, with ca. 10 long setae on basal half of mesial margin ( Fig. 17B View FIGURE 17 ) ................. M. plusiochaeta View in CoL sp. n.

32. Anepisternum nonsetose ....................................................... M. unifurcata Lundström View in CoL

Anepisternum setose ......................................................................... M. meilingae Papp View in CoL