Anthracus philippinensis, Jaeger, 2015

Jaeger, Bernd, 2015, Revision of the maculate species of the Anthracus annamensis group from the East Palaearctic and Oriental Regions. Part 2. A redescription of Anthracus nesophilus (ANDREWES, 1936) and six new species from Nepal, India and SE Asia (Coleoptera, Carabidae, Harpalini, Stenolophina), Linzer biologische Beiträge 47 (2), pp. 1361-1396 : 1366-1367

Anthracus philippinensis nov.sp.

( Figs 7-9 , 35-41 , 80 )

Acupalpus annamensis BATES : ANDREWES 1926: 346

Acupalpus annamensis BATES : ANDREWES 1930: 10 [record from Philippines]

Type material: Holotype: ♂ ( BMNH) labelled "Philippine Is. / Coll. Bottcher / B. M. 1929- 201" and " Philippine Islands / Manila / 14:III:1914 / Coll. Bottcher " [day and month hw]. and "HOLOTYPE ♂ / Anthracus / philippinensis sp. n. / des. B. Jaeger 2015" [red label].

Paratypes: 3♂♂, 4♀♀ ( BMNH) with same labels as the holotype, but 6 specimens additionally with " Acupalpus / anamensis / Bat. / det. Ing. Jedlička ". 2♀♀ ( BMNH) labelled " Philippine Is. / Coll. Bottcher / B. M. 1929-201" [first line yellowish underlined] and " Philippine Islands / Manila / 2:XI:1914 / Coll. Bottcher" and 1♀ additionally with "Acupalpus / anamensis / Bat. / det. Ing. Jedlička ". 5♂♂, 3♀♀ ( SMTD, cJAE) labelled " Luzon / Manila " [yellow label], " 1924/ I " [yellow label] and "Staatl. Museum für / Tierkunde, Dresden", one ♀ additionally with "Anthracus / sp. / det. Dr. E. Schaub."[two first lines hw Schauberger]. 1♂, 3♀♀ ( SMTD) labelled " Luzon, Manila / 10. 1913 / leg. G. Boettcher ", "Samml. O. Langen-/ han. Kauf 1931. 18" and "Staatl. Museum für / Tierkunde, Dresden". 28♂♂, 37♀♀ ( MNB, cJAE) " Philippinen / Manila, lux / leg. Boettcher ". 1♂ ( MNB) with same data, but on backside " 10.5.1914 " [hw]. 49♂♂, 87♀♀ ( MNB, cJAE) labelled " Philippinen / Manila, lux / 2.XI. 1914 " [yellow label]. 1♂ ( RMNH) labelled " Museum Leiden / LUZON, P.I. / Los Baños (Laguna) / campus, a.l. / 24.X. 1964 / J. T. Wiebes " [3rd and 4th line hw] and "Anthracus / nesophilus / ( Andrewes, 1936) / det. B. Jaeger 2013". 1♂, 1♀ ( BMNH) labelled " Los Banos / P. I. Baker ", "Ex Mus. / Coll. Agric. /Phil. Is." and " H.E. Andrewes Coll. / B.M. 1945-97", the ♀ additionally with labels "907" [hw] and "Acupalpus / annamensis / Bates / H.E. Andrewes det." [first three lines hw Andrewes]. 1♂ ( NHMB) labelled " PHILIPPINES, 200 m / SW Panay, 8km E of / Bontol , 10.- 11.Dec / Bolm lgt. 1990". 1♂ ( NHMW) labelled " PHILIPPINEN - Mindoro / E Puerto Galera / Sabang XI. 1992 / leg. Jäch (2)". 1♀ ( NHMB) " PHILIPPINES, 1600 m / Mindanao, 30km W of / MARAMAG , 28.-30. Dec / Bolm lgt. 1990". All paratypes additionally with my label "PARATYPE ♂ or ♀ / Anthracus / philippinensis sp. n. / des. B. Jaeger 2015" [red label].

Additional Material: 4 specimens ( MNB) from the Philippines labelled "Acc.No. 8433 , Lot, Bu. of Sci., P.I." and "Collected by, J. Guerrero ". They were excluded from the type series due to the lack of detailed locality data.

Etymology: The species name refers to the known distribution of the species.

Description: General appearance as figured ( Fig. 7 ). Body length 3.2-3.7 mm (HT 3.4); width 1.3-1.5 mm.

Shiny, pronotum weakly, elytra moderately iridescent. Colour of upper surface usually paler than in A. nesophilus with head and pronotum reddish yellow, but clypeus and labrum paler, and mandibles with inner margins and apices blackish. Rarely, the pronotal disc is weakly infuscated leaving all margins widely reddish yellow. Elytra reddish yellow, with each elytron having a large dark brown central macula, expanding laterally to interval 8 and leaving basal and lateral quarter and first interval reddish yellow. Legs, palpi and first two antennomeres pale yellowish brown, antennomeres 3-11 moderately infuscated. Ventral surface mainly reddish brown, prosternum and epipleura somewhat paler, and pro- and metepisterna weakly infuscated.

Head, pronotum ( Figs 7-9 ) and elytra ( Fig. 7 ) very similar to those of the preceding species with no specific differences in shape or proportions.

Microsculpture of upper surface similar to that of A. nesophilus , but the microsculpture on the frons and anterior half of vertex sometimes almost lacking and on pronotal surface usually less distinct and more lightly impressed.

Metepisterna long and narrowed posteriad, at inner margin about 1.6 times longer than wide at basal margin. Prosternum medially with 5-6 and in front of apical margin with 8 medium long setae (often broken and then insertion points difficult to observe). Prosternal process with 2 distinct setae.

Protarsomeres and mesotarsomeres as in A. nesophilus .

Median lobe of aedeagus ( Figs 35-41 ) similar to that of A. nesophilus , but smaller, somewhat more arcuate ventrad, and internal sac with 3-5 larger, more closely arranged teeth in central part.

Comparisons and remarks: Anthracus philippinensis nov.sp. is the only species of the Anthracus annamensis group so far known from the Philippines and represents a closely related vicariant of A. nesophilus from Java, Sulawesi and Sumbawa. It differs from the latter species by the absolutely smaller, ventrally more clearly curved median lobe of the aedeagus and its internal structures with a group of 3-5 larger, more closely and more centrally arranged teeth (see Figs 35-41 respectively 24-34). Externally both species are very similar and show no significant differences in shape of head, pronotum or elytra. However, in A. philippinensis the pronotum is usually reddish yellow, instead of dark reddish or blackish brown as in A. nesophilus , and the microsculpture on pronotal disc is often less distinctly impressed, but atypical specimens of both taxa show no significant differences in colour or microsculpture.

Distribution A. philippinensis is so far known from the Philippine Islands of Luzon, Mindoro, Mindanao and Panay ( Fig 80 ).

Figures

Fig 80: Distribution: A. nesophilus (ANDREWES) (triangles), A. javaensis nov.sp. (white star), A. sumatraensis nov.sp. (square) A. philippinensis nov.sp. (circle), A. siamensis nov.sp. (black star).

Figs 7-9: Anthracus philippinensis nov.sp. Habitus, head and pronotum. (7) PT, Philippines, Panay, E Bontol, (8) HT, (9) PT, Philippines, Mindoro, E Puerto Galera, Sabang.

Figs 35-41: Anthracus philippinensis nov.sp. Median lobe of aedoeagus, lateral and dorsal aspect. (35, 38) PT, Philippines, Mindoro, E Puerto Galera, (36, 40) PT, Philippines, Panay, E Bontol, (37, 39) PT, Philippines, Luzon, Los Banos.