Racesina oxiana (Boettger, 1889)

Vinarski, Maxim V., Aksenova, Olga V. & Bolotov, Ivan N., 2020, Taxonomic assessment of genetically-delineated species of radicine snails (Mollusca, Gastropoda, Lymnaeidae), Zoosystematics and Evolution 96 (2), pp. 577-608 : 577

24. Racesina oxiana (Boettger, 1889) Figs 8K ; 9J ; Table 3

Limnaeus impurus var. oxiana Boettger 1889: 961, Taf. 27, figs 4, 5.

Cerasina luteola var. oxiana - Zhadin 1952: 177, fig. 79.

Lymnaea (Cerasina) impura - Kruglov and Starobogatov 1993a: 85, fig. 10D, non Troschel 1837 (partim).

Cerasina impura - Vinarski and Kantor 2016: 338, non Troschel 1837.

Cerasina oxiana - Aksenova et al. 2018a: 4.

Racesina oxiana - Vinarski and Bolotov 2018: 332.

TL.

Turkmenistan, middle Amu-Darya River, ‘Tschardschui’ (nowadays Türkmenabat).

Types.

Not traced.

As compared to R. luteola , this snail is characterised by a higher spire and ovate-conical shape of shell. In all other respects, it is conchologically very similar to the type species of the genus. The copulatory apparatus of R. oxiana specimens from Tajikistan, dissected by us, was virtually identical with that of R. luteola from India studied and illustrated by Hubendick (1951). Penis sheath is very narrow, its length is almost equal to length of praeputium or slightly less (see Fig. 8K ).

This species of Racesina is distributed in Central Asia (within the ex-USSR boundaries) and Nepal ( Aksenova et al. 2018a). The oldest available name, based on a type locality situated within this area, is Limnaeus impurus var. oxiana Boettger, 1889. Though Kruglov and Starobogatov (1993b) treated this taxon as identical with Lymnaea impura , the type locality of the latter species is in India, where R. oxiana does not occur. L. impura is, most probably, a junior synonym of R. luteola ( Hubendick 1951; Brandt 1974).

Figures

Figure 8. Copulatory apparatuses of the species in the genera Ampullaceana, Peregriana, Kamtschaticana, Tibetoradix, Racesina, Orientogalba and Bullastra. A. Ampullaceana ampla (11.07.2017, Ukraine, Transcarpathian Region, Irshava River). B. A. balthica (Germany, Mecklenburg-Vorpommern, lake Torgelower See). C. A. dipkunensis (15.09.2016, Ukraine, Transcarpathian Region, Tisa River). D. A. fontinalis (20.08.2017, Russia, Rostov Region, Severskiy Donets River). E. A. lagotis (29.06.2016, Tajikistan, a stream in the Panj River floodplain). F. Peregriana dolgini (13.07.2015, Russia, Tyumen' Region, a floodplain of the Tol'ka oxbow). G. Peregriana peregra (01.07.2006, Russia, Omsk Region, vicinity of Chernoluchye settlement). H. Kamtschaticana kamtschatica (12.04.2014, Russia, Kamchatka Peninsula, Valley of Geysers). I. Tibetoradix hookeri (09 - 10.2012, China, Tibet, a brook, 250 km N of Lhasa). J. T. kozlovi (after Aksyonova et al. 2018 a, modified). K. Racesina oxiana (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village). L. R. siamensis (24.11.2016, Myanmar, a water reservoir on the Yetho River). M. Ampullaceana relicta relicta (2005, Ohrid Lake). N. Orientogalba ollula (08.07.2010, China, Shaanxi Province, a roadside ditch near Yandi village). O. Bullastra cumingiana (Philippines, Luzon Island, Lake Sampaloc). Scale bars: 1 mm. B - after Schniebs et al. (2011), modified.

Figure 9. Shells of species of the genera Ampullaceana, Kamtschaticana, Myxas, Tibetoradix, Racesina, Orientogalba and Bullastra. A. Ampullaceana relicta relicta (05.1975, Macedonia, a spring near St. Naum monastery, Ohrid Lake region; NHMW). B. A. r. pinteri, a paratype (05.06.1973, Macedonia, Prespa Lake near Perovo; MHMW). C, D. Kamtschaticana kamtschatica (Russia, Kamchatka Peninsula, Valley of Geysers; RMBH). E. Myxas glutinosa (14.09.1963, Kazakhstan, floodplain of the Tobol River near Kustanay City; ZISP). F, G. Tibetoradix hookeri, two syntypes (NHMUK). H. T. kozlovi, the holotype (ZISP). I. Racesina luteola (India, Bengal, Paksay, Pabna; NHMUK). J. R. oxiana (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI). K. R. siamensis (" Thailand "; ZMB). L. Orientogalba viridis, a syntype (Guam Island; NMNH). M. O. ollula (1883, Japan, Yokohama; NHMUK). N. Limnaea bowelli, a type (after Preston 1909). O. Orientogalba cf. bowelli (30.06.1924, China, Tibet, Lingka, 14,500 feet a. s. l.; NHMUK). P. Bullastra cumingiana (Philippine, Cebu Island; NHMUK). Scale bars: 2 mm (C, D, F-H, L, M, O), 5 mm (A, B, E, J-K, N, P).