Hecaloidella pacifica, 2021

Xu, Deliang, Webb, Michael D., Shang, Suqin & Zhang, Yalin, 2021, Review of the Pacific leafhopper genus Hecaloidella (Hemiptera: Cicadellidae Deltocephalinae: Selenocephalini) with description of sixteen new species, Zootaxa 4948 (3), pp. 381-403 : 390

H. pacifica Zhang & Webb sp. nov.

Figs. 1E–G , 4K–O

Description. Body length (including tegmina), male: 4.8mm; female: 5.1–5.2mm.

Vertex orange at least distally with a pair of dark brown spots extending from anterior margin to transverse sulcus; face dark brown variably marked with stramineous on clypeus. Thorax dorsally sordid yellow to stramineous. Thorax and abdomen ventrally brown. Forewings brownish hyaline variably marked with yellow on inner area of clavus (see Figs 1E–F ).

Male genitalia with subgenital plate elongate triangular basally with short digitate apex, with a few short fine setae laterobasally. Connective with arms distally touching loop-shaped, stem extended caudally into a long strongly dorsally curved process, with a dorsal medial keel. Aedeagus very small, disassociated from connective; shaft very short, digitate, terminating in a pair of short recurved hook-like processes, with a long stout spine-like process basally; gonopore apical.

Nymph as in Fig. 1G .

Material examined. Holotype: ♂, Indonesia, Sulawesi Utara, Manado, Tomohon, M.R. Wilson, v.-vii.1985, on cloves ( BMNH) . Paratypes: 5♀, same data as holotype including 1♀, NHMUK 010591773 ( Fig. 1E ) ( BMNH) ; 1♀, Malaysia, Sabah, Tawai Plat., 1300ft, south of Telupid , M.E. Bacchus, 8.ix.1977, NHMUK 010591774 ( Fig. 1F ) ( BMNH) .

Other material. Numerous other specimens examined from throughout its range (see checklist) ( AMNH, BMNH, NMNH and NMW).

Etymology. This species takes its name from the widespread distribution in the Pacific.

Remarks. This widespread species on clove, is distinguished by its dark colouration with yellow to orange marking (see Figs 1E–F ). In the male genitalia it is similar to H. nitida but differs in features of the connective and aedeagus (see description).

Figures

FIGURE 1. Habitus images of Hecaloidella species; A, H. bimaculata (holotype); B, H. caudospina (holotype); C, H. nitida (holotype); D, H. nudapenis (holotype); E–G, H. pacifica (paratypes and immature); H, H. borneoensis (holotype); I, H. brevipenis (holotype); J, H. brevispina (holotype); K, H. longipenis (holotype); L, H. quadrispina (holotype); M, H. spinosa (holotype); N, H. brevis (holotype); O, H. brunnifacialis (topotype); P, H. knighti (holotype); Q, H. quadripicta (paratype); R, H. sandakanensis (holotype); S, H. singaporensis (paratype); T, H. brunifasciata (holotype).

FIGURE 4. Male genitalia of H. nitida species group; A–E, H. caudospina; A–B, aedeagus, lateral and caudal view; C–D, connective, lateral and dorsal view; E, style; F–J, H. nudapenis; F–G, aedeagus, lateral and caudal view; H–I, connective, lateral and dorsal view; J, style; K–O, H. pacifica; K–L, aedeagus, lateral and caudal view; M–N, connective, lateral and dorsal view; O, style.