Bromelia pinguin, Linneo, 1753

Espejo-Serna, Adolfo, López-Ferrari, Ana Rosa, Martínez-Correa, Nancy & Pulido-Esparza, Valeria Angélica, 2017, Bromeliad Flora of Chiapas State, Mexico: Richness and Distribution, Phytotaxa 310 (1), pp. 448-450 : 448-450

12. Bromelia pinguin L., Sp. pl. 285. 1753. MAP 3

Piñuela, piñuela de coche, piñuelo, rasca culo, rascaculo (Spanish).

MEXICAN DISTRIBUTION: Chiapas, Colima, Guerrero, Jalisco, Michoacán, Nayarit, Oaxaca, Quintana Roo, Sinaloa, San Luis Potosí, Sonora, Tabasco, Tamaulipas, Veracruz, and Yucatán.

ELEVATION: 0–900 m

SPECIMENS EXAMINED: Acala, R. M. Laughlin 1318 (MICH). Acapetahua, M. C. Escobar O. 722 (CHIP), E. Matuda 16753 (MEXU, MICH), 17721 (CHIPx2, MEXU). Arriaga, A. Shilom Ton 3397 (MICH, US). Benemérito de las Américas, E. Martínez S. 8052 (MEXUx2), 9380 (MEXU, XAL), 10688 (MEXU, XAL). Chiapa de Corzo, E. A. Goldman 1000 ( US). Escuintla, E. Matuda 17877 (en parte) (MEXU). Jiquipilas, O. Farrera S. 297 ( CH, CHIP, HEM, MEXU), J. Santana C. et al. 252 (CHIP, UAMIZ). Marqués de Comillas, G. Castillo et al. 3925 (XAL), E. Martínez S. et al. 24995 (CHAP, MEXU). Ocosingo, G. Aguilar M. & D. Álvarez 1041 (MEXU, UAMIZ), G. Aguilar M. & F. Aguilar 1668 (MEXU, MO, UAMIZ), G. Aguilar M. & G. López A. 8544 (UAMIZx2), J. Calónico S. et al. 23487 (MEXU, UAMIZ), J. Calónico S. et al. 25094 (MEXU, UAMIZ), G. Castillo et al. 4063 (MEXU, XAL), E. Martínez S. 7064 (MEXU), 7752 (MEXU, MO), 8345 (MEXUx2), 9279 (MEXU, MO), 15595 (MEXUx2), A. Reyes-García 2390 (MEXU, MO). Ocozocuautla de Espinosa, C. J. Mazariegos H. 33 (HEM). Pijijiapan, A. Chávez C. 27 (HEM). Totolapa, R. M. Laughlin 858 (ENCB, US). Tuxtla Gutiérrez, B. Cruz S. 17 (HEMx2), E. Palacios E. 1884 (UAMIZx4). Venustiano Carranza, R. M. Laughlin 2889 ( US). Villa Comaltitlán, C. Escobar O. 476 (CHIP). Sin municipio, M. B. Foster & O. Van Hyning 2943 ( US).

CATOPSIS GRISEB.

1. Sepals 10–20 mm long; petals spatulate or lanceolate, 15–25 mm long; capsules (16–)19–24 long, 7–11 mm diameter .... 2

1. Sepals 3–9 mm long; petals triangular to narrowly triangular, 2–12 mm long; capsules 8–15 mm long, 3–7 mm diameter.. ................................................................................................................................................................................................ 3

2. Plants 50–150 cm high; adaxial sepals keeled along its entire length; petals white; rosettes 30-50 cm high ....................... 4

2. Plants 18–45 cm hig; adaxial sepals no keeled or keeled only at the base; petals yellow; rosettes 15-24 cm high .............. 5

3. Leaf sheaths widely elliptic, 5–12 times wider than leaf blades, contracted into a very narrow leaf blade .......................... 7

3. Leaf sheaths elliptic to narrowly elliptic, 1–4(–6) times wider than leaf blades, merging almost imperceptibly to the leaf blade ............................................................................................................................... ........................................................ 8

4. Flowers bisexual; inflorescence simple or rarely once branched .................................................................... C. berteroniana

4. Flowers unisexual; inflorescence twice or very rarely thrice branched ................................................................... C. occulta

5. Floral bracts cucullate, equal or longer than sepals ........................................................................................... C. wangerinii

5. Floral bracts straight at apex, shorter than sepals .................................................................................................................. 6

6. Leaf blades narrowly triangular to lanceolate, attenuate; inflorescence longer than leaves ..................................... C. nutans

6. Leaf blades narrowly oblong, apiculate; inflorescence equal or shorter than leaves .......................................... C. wawranea

7. Leaf blades 2–5 mm wide at its middle portion; sepals green, straw coloured when mature, 4–6 mm long, membranaceous, conspicuously nerved; rosette not bulbous ........................................................................................................... C. juncifolia

7. Leaf blades 10–20 mm wide at its middle portion; sepals yellowish, ochraceous when mature, 6–9(–10) mm long, subcoriaceous, even or nearly so; rosette bulbous ............................................................................................................ C. subulata

8. Peduncle bracts shorter than internodes ................................................................................................................................. 9

8. Peduncle bracts equal or longer than internodes .................................................................................................................. 10

9. Leaves forming a tubular or cylindrical rosette; capsules narrowly ovoid, 8–10 mm long, 3.5–4 mm diameter ...... C. nitida

9. Leaves forming an extended, funnel-shaped rosette; capsules ovoid, (9.5–) 11–15 mm long, (4–) 5–8 mm diameter............ ............................................................................................................................... ............................................. C. sessiliflora

10. Leaf blades 0.8–1 cm wide; upper peduncle bracts narrowly triangular to filiform, 0.2–0.4 cm wide; leaf sheaths 1–2(–3) cm wide ................................................................................................................................................................ C. delicatula

10. Leaf blades 1–8 cm wide; upper peduncle bracts triangular to narrowly triangular, 0.5–3 cm wide; leaf sheaths 3–9 cm

26 • Phytotaxa 310 (1) © 2017 Magnolia Press

ESPEJO-SERNA ET AL.

wide .......................................................................................................................................................................................11

11. Leaf blades narrowly oblong, apiculate ............................................................................................................ C. morreniana

11. Leaf blades narrowly triangular, attenuate. .......................................................................................................................... 12

12. Inflorescence twice branched, 19–33 cm long; primary bracts shorter than sterile basis of the branches; flowers 4–6.5 cm long ............................................................................................................................... ..................................... C. minimiflora

12. Inflorescence once branched, 9–15 cm long; primary bracts longer than sterile basis of the branches; flowers 6–7.5 cm long ............................................................................................................................... ..................................... C. oerstediana