Himallaphus arko, Löbl & Kodada, 2021

Löbl, Ivan & Kodada, Ján, 2021, On the Himalayan Pselaphini (Insecta: Coleoptera: Staphylinidae: Pselaphinae), Biodiversität und Naturausstattung im Himalaya VII, Erfurt: Naturkundemuseum, pp. 349-368 : 353

Himallaphus arko n. sp. ( Figs 4-6 , 11 , 12 )

Holotype ♂: NEPAL: Kosi, ridge S Mangming 2800 m 7.IV.84 LÖbl-Smetana ( MHNG) .

Paratypes: 9 ♀, NEPAL, Koshi Zone, Sankhuawasabha District, with the same data as the holotype ( MHNG) ; 1 ♂, 3 ♀, Koshi Zone, Sankhuawasabha District, Induwa Kola valley. 2800 m, 15.IV.1984, I. LÖbl & A. Smetana ( MHNG) ; 1 ♂, 3 ♀, Koshi Zone, Sankhuawasabha District, “ Bakan ” Wof Tashigaon , 3200 m, 5.IV.1982, A. & Z. Smetana ( MHNG, PCPH) ; 1 ♂, 1 ♀, Koshi Zone, Sankhuawasabha District, above Tashigaon , 3600 m, 6.IV.1982, A. & Z. Smetana ( MHNG) ; 2 ♂, 3 ♀, Koshi Zone, Sankhuawasabha District, above Sheduwa , 3000 m, 31.III-1.IV.1982, A. & Z. Smetana ( MHNG) ; 3 ♂, 3 ♀, Meshi Zone, Panchtar District, above Pahakhola , 2600-2800 m, 31.V.1988, J. Martens & W. Schawaller ( SMNS, MHNG) .

Etymology. The species epithet is a Nepali word meaning next.

Description. Length 1.60-1.75 mm. Headabout 1.4 times as long as wide. Frontal ridges extended to level of eye midlength, narrowed posteriad. Frontal sulcus widened posterior antennal tubercles, parallel-sided, raised at level of eye midlength to form shallow mesal vertexal sulcus reaching level of highest point of vertex. Setose patch in frontal sulcus about 0.07 to 0.09 mm long. Eyes at head mid-length, not prominent, with eight or nine facets. Contours of temples rounded in dorsal view. Lower part of temples explanate, delimited by longitudinal carina, glabrous, very finely pubescent. Upper anterior part of temples with short striae extended onto vertex. Neck and area near neck without striae. Glabrous genal area extended behind eyes and gradually narrowed. Vertex anterior neck convex, with inconspicuous appressed pubescence. Segment IV of maxillary palpus shorter than head, with peduncle much longer than enlarged apical area (ratio 29/16). Length/ width ratio of antennomeres as: I 25/10: II 12/8: III 9/6.5: IV 9/6.5: V 9/6.5: VI 9/6.5: VII 9/6.5: VIII 9/6.5: IX 11/8: X 11/9: XI 30/14. Pronotum hardly longer than wide, strongly convex, somewhat shorter than head, with evenly rounded lateral margins; antebasal sulcus sharply delimited, longitudinal carinae dense, not reaching anterior pronotal margin, striae between carinae narrower than carinae. Anterior 0.02 to 0.03 mm of pronotal disc smooth. Pronotal pubescence moderately short. Elytron gradually impressed toward base, with discal and sutural carinae parallel or converging in anterior two thirds; discal carina wide as orwider than sutural carina; basal elytral margin carinate; anterior third of lateral margin carinate and hardly rounded, pubescence short, similar to pronotal pubescence.

Male characters. Metaventrite with mesal impression delimited by large admesal ridges with inner sides well delimited, diverging apically. Surface of ridges oval, flat and completely covered by patches of short flat setae. Ventrite II with shallow mesal impression. Aedeagus (Figs XY) 0.34-0.35 mm long. Ventral process rather long, slightly bent, broad, at apex nearly as maximal aedeagal width. Parameres gradually widened apically (lateral view). Internal sac with two tufts of long spines.

Measurements of the holotype (in mm). Total length 1.73; length/width of head 0.37/0.26, pronotum 0.34/0.32, elytra 0.40/0.63 (0.22), tergite I 0.55/0.75; length of maxillary palpomere IV 0.48, length of aedeagus 0.27.

Diagnosis. Upper part of temples vertex striate. Contours of temples rounded. Eyes not prominent. Mesal vertexal sulcus extended to level of highest point of vertex. Maxillary palpomere IV with peduncle much longerthan enlarged apical area. Baseof elytra impressed. Ventral process of aedeagus nearly as broad as median lobe. Parameres gradually widened in lateral view. Internal sac with two dense spinose tufts.

Distribution. East Nepal.

Figures

Figs 4-6. Himallaphus arko n. sp.; 4. Habitus. 5. Head and pronotum, dorsal; 6. Head and pronotum in oblique view.

Figs 7-11. Aedeagi in dorsal and lateral views; 7, 8. Nabepselaphus nepalensis n. sp.; 9, 10. Himallaphus pahilo n. sp.; 11. Himallaphus arko n. sp. Scales = 0.1 mm.

Figs 12-16. Aedeagi in dorsal and lateral views; 12. Himallaphus arko n. sp.; 13, 14. Himallaphus bahan n. sp.; 15, 16. Himallaphus chehara n. sp.; Scales = 0.1 mm.