Blackburneus guatemalensis (Bates, 1887)

Dellacasa, Marco, Dellacasa, Giovanni & Gordon, Robert D., 2011, Systematic revision of the American taxa belonging to the genera Alloblackburneus Bordat, 2009, and Blackburneus Schmidt, 1913, with description of seven new species (Coleoptera: Scarabaeidae: Aphodiinae), Insecta Mundi 2011 (204), pp. 1-52 : 24-25

Blackburneus guatemalensis (Bates, 1887)

( Fig. 99-105)

Aphodius guatemalensis Bates, 1887: 88 ; Blackwelder 1944: 212.

Aphodius guatemalensis var. scotinus Bates, 1887: 89 ; Blackwelder 1944: 212.

Aphodius guatemalensis var. chiriquinus Bates, 1887: 89 ; Blackwelder 1944: 212.

Aphodius (Emadiellus) guatemalensis ; Schmidt 1913: 138; Dellacasa 1988: 137.

Aphodius (Emadiellus) guatemalensis var. scotinus ; Schmidt 1913: 138.

Aphodius (Emadiellus) guatemalensis ab. scotinus; Dellacasa 1988: 185.

Aphodius (Emadiellus) guatemalensis var. chiriquinus ; Schmidt 1913: 138.

Aphodius (Emadiellus) guatemalensis ab. chiriquinus; Dellacasa 1988: 108.

Aphodius (Gonaphodiellus) striatipennis Petrovitz, 1962: 111 ; Skelley et al. 2007: 2 (as synonym of A. guatemalensis ).

Blackburneus guatemalensis ; Dellacasa et al. 2002: 193; Skelley et al. 2007: 2.

Blackburneus guatemalensis ab. scotinus; Dellacasa et al. 2002: 197.

Blackburneus guatemalensis ab. chiriquinus; Dellacasa et al. 2002: 200.

Typelocality. Quezaltenango, Guatemala.

Type repository. Natural History Museum, London (type examined).

Redescription. Length 3.0-4.0 mm; oblong, moderately convex, shiny, glabrous. Piceous; head shadowy reddish on each side, pronotum reddish on front margin, elytra brownish, at base with several yellowish strips more orless elongate, shadowy reddish on humeral callus and with a preapical smallyellowish spot ( Fig. 104); sometimes blackish, front angles of pronotum and elytral humeral callus testaceous ( Fig. 103); or testaceous with disc of pronotum, sides and apex of elytra shadowy brownish ( Fig. 102); legs reddish; antennal club brownish. Head with epistome feebly convex, regularly, rather finely, evenly, not closely punctured, punctation somewhat denser and coarser laterally; clypeus subsinuate at middle, round at sides, thinly bordered, edge anteriorlyfaintly upturned; genae round, very shortly ciliate, not protruding fromthe eyes; frontalsuture finely impressed; frontregularly andcoarselypuncturedasepistome. Pronotum transverse, moderately convex, rather sparsely and somewhat irregularly punctured, punctation somewhat denserandcoarseron sides, finerandsuperficialon disc; lateral marginsalmost subparallel, thinly bordered, edge glabrous; hind angles obtuse; base feebly bisinuate. Scutellum elongate, superficially microreticulate, with traces of lateral grooves, sparsely and superficially punctured. Elytra oval, moderately convex, notdenticulateatshoulder, finely striate; striaedistinctly punctured, subcrenulate; interstriae flat, smooth, sparsely very finely and irregularly punctured. Hind tibiae superior apical spur distinctly shorter than first tarsal segment; latter somewhat longer than following three segments combined. Male: metasternal plate distinctly concave; aedeagus Fig. 100-101. Female: metasternal plate nearly flat.

Material examined. COSTA RICA: 19 exx. ( CNCI, DCGI, FSCA, HCGN, MHNG) ; EL SALVADOR: 7 exx. ( DCGI, USNM) ; GUATEMALA: 4 exx. ( DCGI, NYSM, USNM) ; HONDURAS: 2 exx. ( DCGI) ; MEXICO: Chiapas: 213 exx. ( CNCI, DCGI, FSCA, MSNT) ; Colima: 8 exx. ( DCGI, RHTC) ; Distrito Federal: 1 ex. ( NYSM) ; Durango: 5 exx. ( CSCA, NYSM) ; Hidalgo: 1 ex. ( USNM) ; Jalisco: 6 exx. ( CNCI, DCGI) ; México: 4 exx. ( CNCI, DCGI, USNM) ; Michoacán: 66 exx. ( DCGI, MSNT, USNM) ; Morelos: 1 ex. ( DCGI) ; Oaxaca: 99 exx. ( BCBG, CNCI, DCGI, MSNT, NYSM) ; Queretaro: 12 exx. ( CNCI, MSNT) ; San Luis Potosí: 3 exx. ( CNCI, RHTC) ; Veracruz: 3 exx. ( CNCI, DCGI) ; PANAMA: 198 exx. ( CNCI, DCGI, EMEB, FSCA) .

Distribution. Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Colima, Distrito Federal, Durango, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Oaxaca, Queretaro, San Luis Potosí, Veracruz), Panama.

Bionomics. Adults are active all year round and are general surface dung feeders.