Minettia fasciata (Fallén 1820), Fallen, 1820

Semelbauer, Marek & Kozánek, Milan, 2014, The immatures of lauxaniid flies (Diptera: Lauxaniidae) and their taxonomical implications, Zootaxa 3780 (3), pp. 401-454 : 416

Minettia fasciata (Fallén 1820)

Living larvae moderately dorsoventrally flattened, with marked belts and crevices, the distal parts of Malpighian tubules expanded.

Egg. Preparation for SEM was not successful.

First instar ( Figs 65, 66 ). Length 1.38–1.76 mm. Facial mask broad, the peristomal cirri thin and numerous. The ventral organ with two slender digitiform projections. The perstigmatic tufts of posterior spiracles long.

Cephaloskeleton ( Fig. 166 ). Length 0.298 (n=1). Mouth hooks with 5 teeth, basal part hyaline. Epistomal sclerite longer than broad; labrum with two lateral and one apical corner. Ventral bridge arched from ventral view, with rounded cut-out on posterior margin.

Second instar ( Figs 67–70 ). Length 2.20–3.51 mm. Facial mask moderately broad (13–23 scraping cirri per row). The posterior row of peristomal cirri dense, shape variable (from filiform to robust). The anterior spiracle rounded. The first and second thoracic segments smooth except of comb spines; the rest body segments covered by stout spine-like hairs laterally, the posterior-most segments even dorsally.

Cephaloskeleton ( Fig. 181 ). Length 0.58–0.61 mm. Essentially the same as in M. austriaca , subhypostomal sclerites parallel.

Third instar ( Figs 71–76 ). Length 2.82–6.30 mm. Internal side of labial lobe naked. The first thoracic segment smooth. Remaining body segments heavily spinous from dorsal and lateral sides; ventral side with smooth areas. Anterior spiracles fan-shaped, 11 papillae.

Cephaloskeleton ( Fig. 196 ). Length 0.83–0.97 mm. Mouth hooks subparallel. Intermediate sclerite strong, ventral bridge simple and descending posteriad. Epistomal sclerite semilunar with extensive posterior hyaline plate. Subhypostomal sclerite stalked, converging; ligulate sclerites as concave converging plates. Dorsal bridge trapezoid. Dorsal cornua slender; ventral cornua broad from lateral view, medial part of ventral cornua and basal part of vertical plates finely striolated, posterior margin convex and sclerotised. Cibarium slightly pigmented.

Figures

FIGURES 65 – 70. Minettia fasciata, first (65, 66) and second (67 – 70) instars. 65. Cephalic segment, ventral view. 66. Detail of the ventral organ, lateral view. 67. Cephalic segment, ventral view. 68. Anterior spiracle, lateral view. 69. Abdominal segments, lateral view. 70. Last abdominal segment and the anal division, lateral view. A, antenna; A 3 – A 7, third to seventh abdominal segment; AD, anal division; AL, ambulatory lobe; DP, dorsal process; LL, labial lobe; LP, lateral process; MH, mouth hooks MP, maxillary palpus; PC, peristomal cirri; PS, posterior spiracles; SC, scraping cirri; SC 1, first row of scraping cirri; VO, ventral organ.

FIGURES 162 – 167. Cephaloskeletons of first larval instar, dorsal (top), lateral (middle) and ventral (bottom) view. 162. Cnemacantha muscaria. 163. Homoneura biumbrata. 164. H. limnea. 165. Minettia austriaca. 166. M. fasciata. 167. M. flaviventris. c, cibarium; db, dorsal bridge; dc, dorsal cornua; dmh, distal part of mouth hook; es, epistomal sclerite; is, intermediate sclerite; lb, labrum; ls, ligulate sclerite; smh, stalked part of mouth hook; pb, parastomal bar; ss, subhypostomal sclerite; vb, ventral bridge; vc, ventral cornua; vp, vertical plate.

FIGURES 177 – 182. Cephaloskeletons of second larval instar, dorsal (top), lateral (middle) and ventral (bottom) view. 177. Cnemacantha muscaria. 178. Homoneura biumbrata. 179. H. limnea. 180. Minettia austriaca. 181. M. fasciata. 182. M. flaviventris. a, apodeme of ventral cornu c, cibarium; db, dorsal bridge; dc, dorsal cornua; ds, dental sclerite; es, epistomal sclerite; is, intermediate sclerite; lb, labrum; ls, ligulate sclerite; mh, mouth hook; pb, parastomal bar; r, rami; ss, subhypostomal sclerite; vb, ventral bridge; vc, ventral cornua; vp, vertical plate.

FIGURES 71 – 76. Minettia fasciata, third instar. 71. Cephalic segment, ventral view. 72. Detail of the ventral organ. 73. Thoracic segments, dorsal view. 74. Abdominal segments, lateral view. 75. Lateral tubercules of the abdominal segments. 76. Last abdominal segments, and the anal division, dorsal view. A 4 – A 7, fourth to seventh abdominal segments; AD, anal division; DP, dorsal process; LL, labial lobe; LP, lateral process; MH, mouth hooks; PC, peristomal cirri; PS, posterior spiracles; SC, scraping cirri; T 1 – T 3, first to third thoracic segments; VO, ventral organ.

FIGURES 192 – 197. Cephaloskeletons of third larval instar, dorsal (top), lateral (middle) and ventral (bottom) view. 192. Cnemacantha muscaria. 193. Homoneura biumbrata. 194. H. limnea. 195. Minettia austriaca. 196. M. fasciata. 197. M. flaviventris. a, apodeme of ventral cornu; c, cibarium; db, dorsal bridge; dc, dorsal cornua; es, epistomal sclerite; is, intermediate sclerite; lb, labrum; ls, ligulate sclerite; mh, mouth hook; pb, parastomal bar; r, rami; ss, subhypostomal sclerite; vb, ventral bridge; vc, ventral cornua; vp, vertical plate.