Eunotidae new status, 1934

Burks, Roger, Mitroiu, Mircea-Dan, Fusu, Lucian, Heraty, John M., Jansta, Petr, Heydon, Steve, Papilloud, Natalie Dale-Skey, Peters, Ralph S., Tselikh, Ekaterina V., Woolley, James B., van Noort, Simon, Baur, Hannes, Cruaud, Astrid, Darling, Christopher, Haas, Michael, Hanson, Paul, Krogmann, Lars & Rasplus, Jean-Yves, 2022, From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera), Journal of Hymenoptera Research 94, pp. 13-88 : 13

Eunotidae new status

Eunotinae Ashmead, 1904. Type genus: Eunotus Walker, 1834.

Diagnosis.

Antenna with at most 11 flagellomeres. Eyes ventrally divergent. Clypeus with transverse subapical groove. Labrum either exposed and well-sclerotized (most species), or hidden behind clypeus ( Epicopterus Westwood), subrectangular, with marginal setae in a row. Mandibles with 2 or rarely 3 teeth (Fig. 19 ). Subforaminal bridge with postgena separated by lower tentorial bridge. Pronotum transverse. Mesoscutellum with frenum indicated laterally, with axillular sulcus. Mesopleural area without an expanded acropleuron; mesepimeron not extending over anterior margin of metapleuron; only one mesofurcal pit present (Fig. 20 ). All legs with 5 tarsomeres; protibial spur stout and curved; basitarsal comb longitudinal (Fig. 21 ). Metasoma with syntergum, therefore without epipygium.

Discussion.

Eunotidae , as defined herein, was previously known as Eunotini ( Bouček 1988). Moranilidae , composed of species previously included in Eunotinae , differ in having an oblique basitarsal comb that crosses the area where the basitarsal notch would be, in having pits on the mesopleural area of the mesopectus, and in having two mesofurcal pits instead of the single pit usually found in Chalcidoidea . Idioporus affinis ( Idioporidae ) differs in having 4 tarsomeres. Aphelinidae differ in lacking any indication of a frenum, in having a flexible, hidden labrum, usually in having more advanced axilla, and the second phragma extending into the mesosoma. Despite apparent similarities between Aphelinidae and Eunotidae , the two taxa, are seldom confused because Eunotidae have a more strongly sclerotized body, which means that the two families have a very different habitus. Lyciscidae differ in having a subconical pronotum with a median longitudinal carina, whereas the pronotum of Eunotidae is transverse without a visible median carina in dorsal view.

Figures

Figures 19 - 24. 19 - 21 Eunotus sp. (Eunotidae) 19 mandible and labrum in frontal view 20 mesosoma ventral view 21 protibial spur and basitarsal comb 22 Herbertia brasiliensis Ashmead (Herbertiidae) head posterior view 23 Hetreulophus sp. (Hetreulophidae), mesosoma lateral view 24 Zeala walkerae Boucek (Hetreulophidae): propodeum.