Platnickia roble, Grismado & Platnick, 2008

Grismado, Cristian J. & Platnick, Norman I., 2008, Review of the Southern South American Spider Genus Platnickia (Araneae, Zodariidae), American Museum Novitates 3625 (1), pp. 1-20 : 14-16

Platnickia roble , new species

Figures 27, 38, 39, 58, 59

TYPE: Female holotype taken at an elevation of 1100 m at El Roble, Ñuble , Región del Bío-Bío ( VIII) , Chile (Oct. 5–7, 1992; L. Peña), deposited in AMNH.

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Females can easily be distinguished by the short, transverse epigynal median field and the rounded, simple spermathecae (figs. 58, 59).

MALE: Unknown.

FEMALE (holotype, figs. 27, 38, 39): Total length 5.01. Carapace 2.20 long, 1.60 wide. Femur II 1.38 long. Eye sizes and interdistances: AME 0.12, ALE 0.10, PME 0.10, PLE 0.10; AME-AME 0.06, AME-ALE 0.04, PME-PME 0.06, PME-PLE 0.16, ALE-PLE 0.02. Carapace orange brown, with lighter area in front of fovea; legs, endites uniformly orange brown; sternum, chelicerae slightly darker; abdominal dorsum white with cream, oval cardiac band (darkened at middle, with thin), p1-1, r1-1, vp0-1-1(ap), vr1(ap); IV d1-1 (last one very thin), p1-1, r1-1, vp1-1-1(ap), vr1(ap); metatarsi: I p1(ap), v2(ap); II r1(ap), p2(ap), vp0-1-1(ap), vr0-1-1(ap); III v0-2- 2(ap), p0-1-1(subap), dr0-1-1(ap); IV dp1(very thin)-1-1(subap), dr0-1-1, v0-2-2(ap), p1(ap), r1(ap); tarsi: III vr0-0-1-1, vp0-0-1; IV vr0-1-1- 1-1, vp0-0-0-1. Epigynum with short, transverse median field (fig. 58); spermathecae globose, apparently simple (fig. 59).

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Known only from the type locality in central Chile (map 1).

Platnickia wedalen , new species

Figures 28, 29, 40–43, 60–64

Storena bergi (misidentification): Schiapelli and Gerschman de Pikelin, 1974: 85, figs. 15–17 (male only; female is correctly identified).

Map 1. Southern Chile and adjacent Argentina, showing records of Platnickia elegans (squares), P. roble (star), and P. bolson (diamonds).

two white spots), sides, venter cream, except for two paraxial white bands and dark grey mark around spinnerets. Leg spination: femora: I d1-1-1, p1(subap); II d1-1-1; III d1(subap); IV d1(subprox); tibiae: I p1-1, v2(ap); II p11, v2(ap); III d1-1(last one very

TYPES: Male holotype and female allotype (together with one male and three female paratypes) taken at an elevation of 17 m at Chepu , Chiloe´, Región de Los Lagos (X), Chile (Nov. 29, 1981; N. Platnick, R. Schuh) , deposited in AMNH.

ETYMOLOGY: The specific name is from the Mapuche word ‘‘wëdalen’’, meaning ‘‘to be separated or isolated from’’, and refers to the large gap between the two known populations of this species, in the southern Andean forests and on the Falkland Islands.

DIAGNOSIS: Males resemble those of P. bergi in having a membranous tegular apophysis, but in this species the apophysis is slightly longer and runs closely parallel to the ventral surface of the tegular extension, which has a dorsal denticulate keel (figs. 60–62); the retrolateral tibial apophysis has a very acute tip. Females are also very similar to those of P. bergi but have more rounded anterolateral margins on the epigynal median field and more elongated spermathecae (figs. 63, 64). Both sexes show reduced spination on all the legs.

MALE (holotype, figs. 28, 40, 41): Total length 4.28. Carapace 2.14 long, 1.34 wide. Femur II 1.14 long. Eye sizes and interdistances: AME 0.08, ALE 0.06, PME 0.08, PLE 0.06; AME-AME 0.08, AME-ALE 0.02, PME-PME 0.08, PME-PLE 0.12, ALE-PLE 0.06. Carapace red, except frontal, anterolateral areas almost black; chelicerae, endites brown; sternum reddish brown; legs orangish brown, with darkened median bands on tibiae; metatarsi, tarsi slightly darkened; abdominal dorsum dark brown, with two consecutive lateral white bands (posterior pair more elongated, sinuous), sides, venter uniformly dark brown except for few small posterolateral creamy dots. Leg spination: femora: I d1-1- 1(ap); II d1-1-1(ap); III d1(ap); IV d1-0-1(ap); tibiae: I vr1-1-0, p1/0-1/0; II vr1-1-0; III vp1-1- 0-1; IV vp1-1-0-1, r1-1; metatarsi: I v2(ap); II v0-2-2(ap), p1(ap), r1(ap); III vp1-1-0-1(ap), p1(ap), dp0-1/0-1(ap), vr1(ap), r1(ap), dr1(ap); IV dp1(ap), p1(ap), vp1-1-0-1(ap), dr0-1- 1(ap), r0-1-1(ap), vr0-1-1(ap); tarsi: III vr0-0- 0-1, vp0-1; IV vr0-0-1-1-1, vp0-0-0-1-1. Palp as in figs. 60–62.

FEMALE (allotype, figs. 29, 42, 43): Total length 5.33. Carapace 2.30 long, 1.50 wide. Femur II 1.20 long. Eye sizes and interdistances: AME 0.08, ALE 0.10, PME 0.12, PLE 0.08; AME-AME 0.04, AME-ALE 0.04, PME-PME 0.08, PME-PLE 0.18, ALE-PLE 0.04. Carapace brown with reddish cephalic area; sternum, mouthparts dark brown; coxae yellowish brown, remaining podomeres yellowish brown proximally, brown distally (except tarsi entirely brown); abdominal coloration as in male, except posterior dorsal bands replaced by consecutive, irregular, creamy spots, reduced in size toward caudal area; thin creamy lines connect each spot with corresponding spot on opposite side; lateral, ventral small dots more abundant than in male, most ventral dots arranged in paraxial lines; spinnerets light brown. Leg spination: femora: I d1-0-1(all thin); II d1-0-1 (all thin); III d1(subap); IV d1-0-1 (all very thin); tibiae: I vr1(subprox); II vr1(subprox); IV vp1-1-0; metatarsi: I v2(ap); II v2(ap); III p1(ap), vp0- 1-1(ap), vr1(ap), r1(ap), dr1(ap); IV pv1(ap), vp0-1-1(ap), vr0-0-1(ap), r1(ap): tarsi: III vr0- 0-1-1-1, vp0-0-1-0; IV vr0-0-1-1-1, vp0-0-1-1. Epigynum with more or less heart-shaped median field (fig. 63); spermathecae elongat- ed, with medial lobes (heads) larger than laterals (fig. 64).

VARIATION: Some females have two additional ventral spines on metatarsi I and/or II. The form of the white dorsal abdominal marking varies from elongate longitudinal bands to series of consecutive spots.

OTHER MATERIAL EXAMINED: CHILE: Región de la Araucanía (IX): Cautín: Bellavista, N shore Lago Villarrica, Dec. 15–30, 1982, window trap, Valdivian rainforest, elev. 310 m (A. Newton, M. Thayer, AMNH), 1♀; Flor del Lago, 15 km NE Villarrica, Dec. 14, 1984 – Feb. 10, 1985, flight intercept trap, Nothofagus forest, elev. 300 m (S., J. Peck, AMNH), 23. Región de los Lagos (X): Valdivia : 4.1 km W Anticura, Dec. 19–25, 1982, window trap, Valdivian rainforest, elev. 270 m (A. Newton, M. Thayer, AMNH), 23. Llanquihue : 10–14 km E Correntoso, Feb. 3, 1985, Berlese, concentrated moss, disturbed forest, elev. 305 m (N. Platnick, O. Francke, AMNH), 13, 2♀; 7 km N Ensenada, Nov. 26, 1981, concentrated moss, litter, elev. 400 m (N. Platnick, R. Schuh, AMNH), 1♀. Chiloe´: 5 km N Quellón, Dec. 1, 1981, elev. 107 m (N. Platnick, R. Schuh, AMNH), 1♀. Palena: 25–27 km N Chaitén, Jan. 17, 1986, Berlese, moss, wet virgin forest, elev. 40 m (N. Platnick, P. Goloboff, R. Schuh, AMNH), 2♀. Región Aisén (XI): Aisén: 30 km N Puyuhuapi, Jan. 29, 1985, sifted moss on logs, elev. 100 m (S., J. Peck, AMNH), 1♀. ARGENTINA: Río Negro: Parque Nacional Nahuel Huapi, Puerto Blest, Jan. 7–20, 2000, sifting humus (L. Lopardo, A. Quaglino, MACN), 1♀. FALKLAND ISLANDS: East Falkland: Murrell Valley, Dec. 3, 1974 (S. Coscarón, AMNH), 13; ‘‘Fitz Roy Stanley’’, Feb. 1971 (M. Rumboll, MACN 6669), 13 (not ♀).

DISTRIBUTION: Chile (Cautín to Aisén), Argentina (Río Negro), and the Falkland Islands (where the species is apparently sympatric with P. bergi , map 2).