Asaphesinae incertae sedis new placement, 2000

Burks, Roger, Mitroiu, Mircea-Dan, Fusu, Lucian, Heraty, John M., Jansta, Petr, Heydon, Steve, Papilloud, Natalie Dale-Skey, Peters, Ralph S., Tselikh, Ekaterina V., Woolley, James B., van Noort, Simon, Baur, Hannes, Cruaud, Astrid, Darling, Christopher, Haas, Michael, Hanson, Paul, Krogmann, Lars & Rasplus, Jean-Yves, 2022, From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera), Journal of Hymenoptera Research 94, pp. 13-88 : 13

Asaphesinae incertae sedis new placement

Asaphinae Ashmead, 1904. Type genus: Asaphes Walker, 1834. Junior homonym of Asaphidae Burmeister, 1843.

Asaphesinae Burks & Heraty, 2020, replacement name.

Diagnosis.

Antenna with 12 flagellomeres, including a small 4th clavomere. Clypeus with transverse subapical groove. Head dorsally with temple separating posterior margin of eye from that of the head. Labrum exposed, well-sclerotized, subrectangular with marginal setae in a row. Mandibles with 2 or 3 teeth. Subforaminal bridge with postgena separated by lower tentorial bridge; occipital carina present. Mesoscutellum with frenum indicated at least laterally, and with axillular sulcus. Mesopleural area without an expanded acropleuron, with pits; mesepimeron not extending over anterior margin of metapleuron; two mesofurcal pits usually present (Fig. 59 ) (absent in the fossil genus Coriotela ). All legs with 5 tarsomeres; protibial spur stout and curved; basitarsal comb oblique. Metasoma with syntergum, therefore without epipygium.

Discussion.

The scope of Asaphesinae is much reduced with the removal of Enoggerinae new subfamily to a separate incertae sedis subfamily in Chalcidoidea , and Bairamlia Waterston to Sphegigastrini ( Pteromalidae ). Asaphesinae is part of a set of taxa with two mesofurcal pits and some other shared features, but which do not form a clade in molecular analyses (Cruaud et al, submitted), including Enoggerinae , Herbertiidae , and Moranilidae . Asaphesinae differ from Herbertiidae and Moranilidae in having 12 flagellomeres instead of a maximum of 10. Enoggerinae differ in lacking a temple, thus with the posterior margin of the eye dorsally meeting that of the head. Some Asaphesinae resemble Pteromalidae in habitus, differing in the clypeus, reduced mandibles, labrum, oblique basitarsal comb, and presence of two mesofurcal pits, but also having features that are rarely found in Pteromalidae , such as an occipital carina and the mesopleural area with pits.

Figures

Figures 55 - 60. 55 Erotolepsia sp. (Spalangiidae, Erotolepsiinae): mesosoma and base of metasoma dorsal view 56 a, 57 Systasis sp. (Systasidae) 56 a apex of clypeus without subapical groove 57 mesosoma ventral, mesotrochantinal plate and mesofurcal pit 58 Trisecodes africanum Gumovsky (Pirenidae, Trisecodinae): antenna 56 b, 59 Asaphes sp. (Asaphesinae, incertae sedis) 56 b clypeal subapical groove 59 mesosoma ventral 60 Austrosystasis atricorpus Girault (Austrosystasinae, incertae sedis) 60 head frontal view, mesosoma lateral view.