Tylopus veliger, Likhitrakarn, Natdanai, Golovatch, Sergei I., Prateepasen, Rujiporn & Panha, Somsak, 2010

Likhitrakarn, Natdanai, Golovatch, Sergei I., Prateepasen, Rujiporn & Panha, Somsak, 2010, Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae), ZooKeys 72, pp. 23-68 : 30-32

Tylopus veliger sp. n. Figs 1012

Holotype

♂ (CUMZ), Thailand, Nan Province, Pua District, Ton Tong Waterfall, ca 1130 m, 19°10'52N, 101°5'45E, 10.10.2009, leg. S. Panha, J. Sutcharit & N. Likhitrakarn.

Name:

To emphasize the velum-shaped end of gonopod lobe l.

Diagnosis:

Differs from congeners except Tylopus perplexus Golovatch and Enghoff 1993 in the distal part of gonopod lobe l being velum-shaped and supplied with two denticles, from Tylopus perplexus in the gonopod lacking spines m and q, as well as in a much shorter and knife-shaped spine z, and a rudimentary spine h.

Description:

Length ca 28 mm, width of midbody pro- and metazona 2.0 and 2.6 mm, respectively. Coloration of live animal and alcohol material rather uniformly dark brown to blackish (Fig. 1A); calluses of paraterga only slightly flavous, brown; antennomeres 1-6 and genae light brown; venter and legs contrastingly yellowish to light brown (Fig. 10 A–G).

All characters as in Tylopus bispinosus sp. n., except as follows.

Clypeolabral region of head very densely setose, but vertigial region bare. Antennae short and barely reaching behind segment 2 dorsally. In width, head = segments 3 and 4 <collum <segment 2 <5-16; thereafter body gradually and gently tapering towards telson (Fig. 10 A–G). Collum with small, narrowly delimited, rounded, strip-shaped paraterga (Fig 10B, C).

Metaterga with two transverse rows of long setae: 2+2 in anterior and 2(3)+2(3) in posterior row, the latter often abraded, but then readily traceable as insertion points. Axial line thin, in places incomplete, but readily visible on both halves of metaterga. Paraterga strongly developed (Fig. 10 A–G), lying relatively low (at 1/2-1/3 midbody height), evidently inclined ventrolaterally, pointed caudally and acutangular already from segment 2, especially strongly so and surpassing rear tergal contour on segments 16-19; calluses slightly thinner on poreless segments than on pore-bearing ones; poreless calluses with two lateral setigerous incisions, but with only a single, more evident one (anterior) on pore-bearing calluses (Fig. 10 B–G). Transverse sulcus evident on metaterga 5-18, reaching bases of paraterga, evident and rather deep, finely, densely and clearly ribbed at bottom. Stricture between pro- and metazona very clearly ribbed (Fig. 10 B–G). Epiproct tip clearly emarginate, pre-apical papillae evident (Fig. 10 F–H). Hypoproct semi-circular, caudal setae strongly separated (Fig. 10H). Pleurosternal carinae as complete ridges on segments 2-4, thereafter broken into an anterior bulge and a caudal tooth, both growing increasingly reduced until segment 16 (Fig. 10C, E, F).

Sterna rather densely setose, without modifications except for a subquadrate, setose, sternal lobe between coxae 4 (Fig. 10I, J). Legs relatively short, ca 1.2-1.3 times as long as midbody height, evidently incrassate (Fig. 10C, F, K); prefemora distinctly bulged laterally and clothed with mostly adpressed setae ventrally (Fig. 10K), acropodites likewise with very dense, mostly adpressed setae ventrally; postfemora and tibiae slightly bulged ventrally; tarsal brushes missing.

Gonopods (Figs 11, 12) with lobe l well-demarcated, high and prominent, apically with a pointed fan-shaped structure (= velum) and two denticles; spine h very small, dentiform; spine z prominent, knife-shaped, lying above l on lateral side.