Pseudochactas ovchinnikovi, Gromov 1998 LP, 2303

Loria, Stephanie F., Ehrenthal, Valentin L., Nguyen, Anh D. & Prendini, Lorenzo, 2022, Climate Relicts: Asian Scorpion Family Pseudochactidae Survived Miocene Aridification in Caves of the Annamite Mountains, Insect Systematics and Diversity 6 (6), pp. 1-21 : 9

P. ovchinnikovi (68%), Pseudochactas (58%), the Thiên Ðu lation of V. canhi (28%), and the Southeast Asian clade comprising Troglokhammouaninae Prendini et al. 2021 and Vietbocapinae. Jackknife support values in the separate analysis of the morphological dataset with parsimony and equal weighting were also high (≥ 79%) except for P. ovchinnikovi (70%), A. lao (66%), the Thiên Ðu population of V. canhi (54%) and intraspecific relationships among A. lao , P. ovchinnikovi , T. steineri and V. canhi (> 10%).

Simultaneous analyses of the morphological and molecular datasets with BI, ML and parsimony with equal or implied weighting (k = 1, 3, 10, 60, 100) were congruent with separate analyses of the molecular and morphological datasets and recovered all subfamilies, genera, species and populations as monophyletic. Posterior probability and bootstrap support values obtained by simultaneous analysis of the morphological and molecular datasets were generally high in the analyses with BI (≥0.99), ML (≥99%), and parsimony with equal weighting (≥ 100%). Jackknife support values were also high (≥83%) in the parsimony analysis with equal weighting except for the following, which received low to moderate support: P. ovchinnikovi (63%) and intraspecific relationships among V. canhi and T. steineri (>10%). Unambiguous morphological synapomorphies were optimized on the ML tree obtained by simultaneous analysis of the morphological and molecular datasets, the preferred phylogeny ( Fig. 2 ). Output files from phylogenetic analyses are publicly available in figshare: 10.6084/m9.figshare.21067528.

Figures

Fig. 2. Preferred phylogeny (A) of the relictual Asian scorpion family Pseudochactidae Gromov 1998 based on simultaneous analysis of the morphological and molecular datasets using Maximum Likelihood with unambiguous morphological synapomorphies optimized. Black circles indicate uniquely derived apomorphic states, white circles, parallel deviations of apomorphic states. Numbers above circles indicate characters, numbers below indicate states (Supp Appendices 2 and 3 [online only]). Dorsal habitus (B–F) of five species of Pseudochactidae: Pseudochactas ovchinnikovi Gromov 1998, Babatag, Uzbekistan, ♀ (AMNH) (B), Troglokhammouanus steineri Lourenço 2007,Tham Xe Bang Fai, Laos, ♀ (AMNH) (C), Aemngvantom thamnongpaseuam Prendini et al. 2021,Tham Nong Pa Seuam,Laos, holotype ♀ (AMNH) (D), Aemngvantom lao (Lourenço 2012),Tham Nam Lot (Lod), Laos, ♀ (AMNH) (E) and Vietbocap canhi Lourenço and Pham 2010,Tiên Sơn, Vietnam,♀ (AMNH) (F). Scale bars = 10 mm.