Glyphiulus impletus Jiang, Guo, Chen & Xie, 2018

Zhao, Yi, Guo, Wan-Ru, Golovatch, Sergei I. & Liu, Wei-Xin, 2022, Revision of the javanicus species group of the millipede genus Glyphiulus Gervais, 1847, with descriptions of five new species from China (Diplopoda, Spirostreptida, Cambalopsidae), ZooKeys 1108, pp. 89-118 : 89

Glyphiulus impletus Jiang, Guo, Chen & Xie, 2018

Glyphiulus impletus Jiang, Guo, Chen & Xie, 2018: 171.

Material examined.

9 ♂ (SCAU WL38), China, Guangxi Zhuang Autonomous Region, Hechi City , Fengshan County, Jiangzhou Underground Corridor , 24°30'4"N, 106°53'46"E, 850 m alt., 2013-VI-30, leg. Tian Mingyi, Liu Weixin, Lin Wei, Yin Haomin & Huang Sunbin. 4 ♂, 2 ♀ (SCAU WL39), China, Guangxi Zhuang Autonomous Region, Baise City , Linyun County, Cave Fengniu Dong , 24°28'39"N, 106°37'52"E, 700 m alt., 2017-VI-9, leg. Tian Mingyi, Liu Weixin, Wang Xinhui & Tang Mingruo.

Remarks.

This species has been found to have a relatively wide distribution in Guangxi, involving the Lingyun, Fengshan, Nandan and Donglan Counties. The new samples add only the records of two cave collections (Fig. 16 ). The above material is in good agreement with the original description by Jiang et al. (2018), while intraspecific p-distances are 2.4-8.2%, based on DNA-barcoding. Based on molecular evidence, G. impletus differs from all other Glyphiulus species analysed from between 11.9% (compared to G. calceus ) and 24.9% (compared to G. foetidus ).

Figures

Figure 16. A distribution map of the Glyphiulus formosus species group in China. 1 G. recticullus 2 G. foetidus 3 G. impletus 4 G. calceus 5 G. fortis 6 G. portaliformis Zhao & Liu, sp. nov. 7 G. pulcher 8 G. scutatus Zhao & Liu, sp. nov. 9 G. echinoides 10 G. xiniudong Zhao & Liu, sp. nov. 11 G. sinuatoprocessus Zhao & Liu, sp. nov. 12 G. conuliformis Zhao & Liu, sp. nov. 13 G. formosus 14 G. hainanensis. Abbreviations: FJ: Fujian, HN: Hainan, HuN: Hunan, GD: Guangdong, GX: Guangxi, GZ: Guizhou, ZJ: Zhejiang.