Cyrtorhininae Guinot, 1993

Van Bakel, Barry W. M., Guinot, Danièle, Artal, Pedro, Fraaije, René H. B. & Jagt, John W. M., 2012, A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata) 3215, Zootaxa 3215 (1), pp. 1-216 : 107-108

Subfamily Cyrtorhininae Guinot, 1993

Cyrtorhininae Guinot, 1993b: 1330 .

Type genus. Cyrtorhina Monod, 1956 .

Genera included. Antonioranina n. gen.; Cyrtorhina Monod, 1956 .

Diagnosis. Carapace ovate, variously wide, convex in both directions. Cervical groove obsolete, branchiocardiac groove indistinct. Posterolateral margins straight, anterolateral margins rounded; margins rimmed or lined by granules. Anterolateral margin with 2, 3 teeth, subhepatic spine may be present. Cuticle microstructure diverse: upright nodes, fungiform nodes, inclined nodes, pits.

Front short, widely triangular, weakly to strongly sulcate, with sharp, anteriorly directed tooth on each side. Supraorbital margin with 2 fissures, inner fissure proportionally short, outer fissure longer, separating extraorbital teeth; outer extraorbital tooth robust, falciform. Orbits small; eyestalk short, of 3 elements: proximal article reduced, only ventrally visible; second article triangular, elongated; third dorsally exposed, calcified, strongly arched, with fungiform granules; cornea small. Antennary fossae situated posterior to antennulary fossae. Antennule deeply inserted, completely hidden by antenna, basal article notched distally, following articles much smaller; flagellum short. Proepistome distinct. Antennae massive, meeting medially, with external surface granular; articles 1, 2 fused, urinal opening at inner angle; article 3 expanded, with spatulate dorsal lobe, pointed ventral lobe; article 4 with salient anterodorsal angle; article 5 short; flagellum developed. Epistome triangular, medially raised, keeled. Subantennary lobe of pterygostome not delimited.

Mxp3 operculiform; endopod long, blunt anteriorly, merus shorter than ischium; palp short, concealed. Exopod short, wider than endopod, anteriorly pointed. Pterygostome granular, ridged, with rows of dense setae, completely setiferous posteriorly. Exposed pleurites 5–7 forming large, unexcavated plate; branchiostegite developed, high, overhanging exposed pleurites, limited by P5 apposed along branchiostegal margins. Sternum/pterygostome junction very narrow, recessed. Sternum/exposed peurite connections large between P1, P2, narrower between P2, P3.

Thoracic sternum narrow all along, thus P1‒P5 coxae approximated. A strong deflection at level of sternite 8. Sternite 1, 2 in a lower level, concealed; sternite 3 exposed, as small, V-shaped, horizontally extended strip; sternite 4 narrow, not laterally expanded except the slender, recessed anterolateral extensions forming sternum/pterygostome junction; sternites 4, 5 expanded laterally between P1, P2; sternites 6, 7 conspicuously narrow, linear; sternite 8 perpendicular to sternite 7. Medial line along sternites 5–8. Spermathecal apertures recessed, small, contiguous.

Chelipeds homochelous, homodontous. Basis-ischium short, immoveably fused with massive merus. Propodus longer than wide, narrow, upper, lower margins smooth, except for acute superodistal spine; fingers long, acicular, crossing at acute tips; dactylus deflected, proximally with long acute spine, prehensile border unarmed; fixed finger only slightly deflected, prehensile border with 3 acute spines proximally. P2–P4 moderately stout; carpus short, modified. P2–P4 dactylus spatulate on P2, sickle shaped on P3, P4. P5 dorsal, moderately reduced, granular, lying along posterolateral carapace margin, adapted to branchiostegal edge; dactyl narrow, elongated.

Abdomen proportionally small but rather wide in male, freely articulated, 6 articles plus telson, incompletely folded, first 4 articles dorsal, first somite as wide as posterior carapace margin. Female abdomen only slightly enlarged, thus sexual dimorphism indistinct. Uropods or sockets absent.

Large posterior branchial orifices present.