Electropsychops handlirschi, Lu, Xiumei, Zhang, Weiwei, Ohl, Michael & Liu, Xingyue, 2017

Lu, Xiumei, Zhang, Weiwei, Ohl, Michael & Liu, Xingyue, 2017, New genus and species of silky lacewing (Insecta: Neuroptera: Psychopsidae) from the mid-Cretaceous Burmese amber, Zootaxa 4291 (2), pp. 373-383 : 376-379

Electropsychops handlirschi sp. nov.

( FigS. 1–4 )

Diagnosis. AS fOr the genuS.

Description. Adult male bOdy length 6.79 mm; head 0.75 mm lOng; diStance between cOmpOund eyeS 0.80 mm; prOthOrax 0.38 mm lOng and 0.61 mm wide; meSOthOrax 0.96 mm lOng and 0.97 mm wide; metathOrax 0.90 mm lOng and 0.93 mm wide; preServed part Of left fOrewing 6.52 mm lOng and 4.39 mm wide; preServed part Of right fOrewing 8.18 mm lOng and 3.80 mm wide; left hind wing 8.02 mm lOng and 3.82 mm wide; abdOmen length ca. 4.42 mm lOng; genitalia 1.23 mm lOng.

Head OrthOgnathOuS, with vertex diStinctly dOmed. COmpOund eyeS large, Semi-glObular; OcelluS abSent. Antennae nOt preServed ( FigS. 1 , 3 A).

PrOthOrax Slightly lOnger but narrOwer than head, with lOng and SparSe Setae; meSOthOrax rObuSt, brOader than prO- and metathOrax. LegS Slightly StOut; Only partS Of fOre- and midleg viSible in dOrSal view, with ShOrt Setae; twO tibial SpurS preSent; tarSi 5-Segmented, tarSOmere 1 aS lOng aS cOmbinatiOn Of tarSOmere 2 and 3, tarSOmere 2 Slightly ShOrter than tarSOmere 3, almOSt aS lOng aS tarSOmere 4, tarSOmere 5 ShOrteSt; Simple pretarSal clawS paired, ShOrt, StrOnly curved.

FOrewing ( FigS. 1 , 2 A–B, 3D–E): ElOngate and OvOid; tranSparent and immaculate; trichOSOrS preSent alOng entire wing margin; Only the baSal nygma preSent; humeral plate rOund; humeral veinlet recurrent, with Seven brancheS; cOStal Space brOad prOximally, but Slightly narrOwed diStad, with 44 cOStal crOSSveinS, mOSt Of which Simple On prOximal and median part Of wing except fOr thOSe On pterOStigmal regiOn mOStly fOrked diStad; nO interlink veinletS; ScA preSent, ShOrt, diStally running tOward ScP; ScP rather StOut prOximally, diStally Slightly curved pOSteriad; SubcOStal Space Slightly wider than RA Space, with nine crOSSveinS On prOximal half and diStally with a ShOrt crOSSvein cOnnecting ScP and RA; RA Straight prOximally and Slightly SigmOid diStally, with Only One marginal fOrk; RA Space with 9–10 crOSSveinS On prOximal half and diStally with a ShOrt crOSSvein, which iS almOSt cOntinuOuS with diStal Scp-ra crOSSvein, fOrming a vena triplica; RP+MA Originated frOm RA near wing baSe, RP with 12 brancheS, Several RP brancheS deeply fOrked; MA Simple alOng almOSt entire length in left fOrewing, but deeply fOrked intO twO main brancheS in right fOrewing; MP Originated frOm R, and fOrked Slightly prOximad itS midlength intO twO main brancheS, which have Only quite a few marginal fOrkS; CuA and CuP diverging near wing baSe; CuA pectinately branched frOm abOut itS midlength; CuP deeply and dichOtOmOuSly branched; A1 pectinately and prOfuSely branched, fOrming a triangular regiOn; A2 ShOrt, with a few brancheS; a cOmplete Outer gradate SerieS Of crOSSveinS preSent On branching area Of RP+MA.

Hind wing ( FigS. 1 , 2 C): Smaller than fOrewing, diStinctly brOadened diStad, with anterOdiStal cOrner Slightly prOduced; tranSparent and immaculate; trichOSOrS preSent alOng entire wing margin; nygma abSent; humeral veinlet recurrent, with fOur Simple brancheS; cOStal Space brOad thrOughOut, width in pterOStigmal regiOn abOut fOur timeS the cOmbined width Of adjacent ScP and RA SpaceS, with 35 cOStal crOSSveinS, mOSt Of which are Simple On prOximal and median part Of wing except fOr thOSe On pterOStigmal regiOn mOStly fOrked diStad; nO interlink veinletS; ScP diStally Slightly curved pOSterad and terminally bending tOward RA by a ShOrt veinlet; SubcOStal Space Slightly narrOwer than RA Space, SubcOStal crOSSvein undetected; RA Straight, with Only One marginal fOrk; RA Space with Six crOSSveinS On prOximal half and diStally with a ShOrt crOSSvein, which iS almOSt cOntinuOuS with terminal veinlet Of ScP, fOrming vena triplica; RP+MA Originated frOm RA near wing baSe, RP with 13 brancheS, Several RP brancheS deeply fOrked; MA Simple, with Stem preSent aS a SigmOid, Oblique veinlet; MP fOrked Slightly prOximad itS midlength intO twO main brancheS, anteriOr One Of which bearS a marginal fOrk; CuA and CuP diverged near wing baSe; CuA pectinately branched frOm abOut itS midlength intO 14 Simple brancheS; CuP Only trifurcated frOm itS midlength; A1 pectinately and prOfuSely branched, fOrming a Small triangular regiOn; A2 ShOrt, with a few brancheS; a cOmplete Outer gradate SerieS Of crOSSveinS preSent On branching area Of RP+MA, arcuately arranged.

AbdOmen Slenderly elOngate. Male genitalia diScernible Only in dOrSal view, cOvered with lOng Setae; tergum 9 Subquadrate, nearly twice aS wide aS lOng, with rOunded pOSterOlateral cOrnerS; ectOprOctS paired and elOngate, each ectOprOct prOximally flat and brOad, laterally with SemiSpherical calluS cercuS, remaining diStal part digitifOrm and Slightly incurved; Subanale preSent and OvOid ( FigS. 1 , 4 ).

Type material. HOlOtype: EMTG BU-001914: amber piece with an almOSt cOmplete male Of Electropsychops handlirschi sp. nov. (diStal partS Of fOrewingS nOt preServed; right hind wing largely fOlded under right fOrewing), a hymenOpteran and twO dipteranS; it iS pOliShed in the fOrm Of arched pentagOn cabOchOn, clear and tranSparent, with length × width abOut 20.1 × 12.5 mm, height abOut 6.0 mm.

Etymology. The new SpecieS iS dedicated tO the late Dr. AntOn HandlirSch, whO eStabliShed the family PSychOpSidae .

Figures

FIGURE 1. Habitus photograph of Electropsychops handlirschi gen. et sp. nov., holotype male, dorsal review. Scale bar: 1 mm.

FIGURE 2. Electropsychops handlirschi gen. et sp. nov., holotype male. A. Drawing of right forewing; B. Drawing of left forewing; C. Drawing left hind wing. Scale bar: 1 mm.

FIGURE 3. Electropsychops handlirschi gen. et sp. nov., holotype male. A. Photograph of head; B. Photograph of right foreleg; C. Drawing of right foreleg; D. Photograph of left forewing base; E. Photograph of right forewing base. ts: tibial spurs. Scale bar: 0.5 mm (A – B, D – E); 0.1 mm (C)

FIGURE 4. Electropsychops handlirschi gen. et sp. nov., holotype male. A. Photograph of genitalia; B. Drawing of genitalia. T: tergum; c: callus cercus; e: ectoproct; sa: subanale. Scale bar: 0.2 mm.