Eufriesea Cockerell, 1908

Nemésio, André & Rasmussen, Claus, 2011, Nomenclatural issues in the orchid bees (Hymenoptera: Apidae: Euglossina) and an updated catalogue, Zootaxa 3006, pp. 1-42 : 19-23

Genus Eufriesea Cockerell, 1908

Plusia Hoffmannsegg, 1817: 52 –53 (not Ochsenheimer, 1816; Lepidoptera ). Type species: Plusia superba Hoffmannsegg, 1817 , monobasic.

Eumorpha Friese, 1899: 126 (not Hübner, 1807; Lepidoptera ). Type species: Euglossa pulchra Smith, 1854 , by designation of Cockerell, 1908: 41.

Eufriesea Cockerell, 1908: 41 , replacement for Eumorpha Friese, 1899 . Type species: Euglossa pulchra Smith, 1854 , autobasic (see Eumorpha Friese, 1899 ).

Eufriesia Lutz & Cockerell, 1920: 544, unjustified emendation of Eufriesea Cockerell, 1908 .

Euplusia Moure, 1943: 189 –190, replacement for Plusia Hoffmannsegg, 1817 . Type species: Plusia superba Hoffmannsegg, 1817 , autobasic (see Plusia Hoffmannsegg, 1817 ).

Paleoeuglossa Poinar, 1998: 30. Type species: Paleoeuglossa melissiflora Poinar, 1998 , by original designation. (According to D.W. Roubik, not an Eufriesea due to the presence of scutellar tuft).

Eufriesea aeneiventris [ Euglossa ] ( Mocsáry, 1896: 5–6), image: Nemésio, 2009a: 35 a–f, Lectotype Ƥ, designated by Moure, 1967b: 406, HNHM: BRAZIL, Espírito Santo.

Eufriesea andina [ Euglossa (Eumorpha) ] ( Friese, 1925: 27–28), Lectotype Ƥ, designated by Moure, 1967b: 406, ZMHB: PERU, Cusco, Vilcanota.

Eufriesea anisochlora [ Euplusia ] ( Kimsey, 1977: 8–10, figs. 10, 22, 34), Holotype 3, UCDC: PANAMA, Panama, Cerro Jefe, leg. Dressler.

Eufriesea atlantica Nemésio, 2008 (“2007”): 148–149, image: Nemésio, 2009a: 40 a–f, Holotype 3, UFMG: BRAZIL, Minas Gerais, Marliéria, Parque Estadual do Rio Doce, leg. Nemésio.

Eufriesea auriceps [ Euglossa (Eulema!)] ( Friese, 1899: 156), Holotype 3, ZMHB: PARAGUAY, Central, Asunción.

danielis [ Eumorpha combinata ssp.] ( Schrottky, 1907: 55, 56–57), image: Nemésio, 2009a: 16a–f (false type), Holotype Ƥ,?; MPSP (=false type): PARAGUAY, Central, Asunción, leg. Anisits: Not synonymized in Moure et al. (2007).

nordestina [ Euplusia ] ( Moure, 1999: 100–101), image: Nemésio, 2009a: 21a–f, Holotype Ƥ, DZUP: BRAZIL, Paraíba, Santa Luzia, 300 m, leg. Portela: Not synonymized in Moure et al. (2007).

aridicola [ Euplusia ] ( Moure, Neves, & Viana, 2001: 842–844, figs. 1–4), image: Nemésio, 2009a: 20a–f, Holotype 3, DZUP: BRAZIL, Bahia, Barra, leg. Neves & Viana: Not synonymized in Moure et al. (2007).

Eufriesea auripes [ Euglossa (Eulema!)] ( Gribodo, 1882: 266–267), Holotype 3, MSNG: FRENCH GUIANA, Cayenne.

Eufriesea bare González & Gaiani, 1989: 151 –152, fig. 2, Holotype 3, MIZA: VENEZUELA, Amazonas, San Carlos de Río Negro, leg. Chacón & Yépez.

Eufriesea boharti [ Euplusia ] ( Kimsey, 1977: 10, figs. 11, 23, 35), Holotype 3, UCDC: VENEZUELA, Aragua, Rancho Grande, leg. Bohart.

Eufriesea brasilianorum [ Euglossa (Eumorpha) combinata ssp.] ( Friese, 1899: 147), image: Nemésio, 2009a: 43 a–b, 45a–d, Holotype Ƥ, ZMHB: BRAZIL, Espírito Santo.

Eufriesea buchwaldi [ Euglossa (Eufriesea) ] ( Friese, 1923: 27–28), Lectotype Ƥ, designated by Moure, 1967b: 407, ZMHB: PERU.

Eufriesea chaconi González & Gaiani, 1989: 149 –150, fig. 1, Holotype 3, MIZA: VENEZUELA, Bolívar, La Paragua, E of Chigua river, 425m, leg. Instituto de Zoología Agrícola Expedition.

Eufriesea chalybaea [ Euglossa (Eufriesea) ] ( Friese, 1923: 27), Lectotype 3, designated by Moure, 1967b: 407 (cf. ICZN Art. 74.6), AMNH (type status unconfirmed): BOLIVIA, Cochabamba, Tarata.

Eufriesea chrysopyga [ Euglossa ] ( Mocsáry, 1898: 497–498), Holotype; invalid lectotype designation of holotype Ƥ, Moure, 1967b: 407, HNHM: BOLIVIA, La Paz, Cordillera Real, Zongo (“Songo”).

Eufriesea coerulescens [ Euglossa ] (Lepeletier de Saint Fargeau, 1841: 11), Lectotype 3, designated by Moure, 1967b: 407 (cf. ICZN Art. 74.6), MNHN: MEXICO: Original spelling appears to be coerulescens , not caerulescens as currently used.

simillima [ Euplusia ] (Moure & Michener in Moure, 1965: 275–277), Holotype 3, SEMC: MEXICO, Chihuahua, Maguarichi, leg. Wood & Karren.

Eufriesea combinata [ Euglossa ] ( Mocsáry, 1897: 446), image: Nemésio, 2009a: 14 a–f, Holotype; invalid lectotype designation of holotype Ƥ, Moure, 1967b: 407, HNHM: BOLIVIA, Potosí, San Antonio.

Eufriesea concava [ Euglossa (Eulema!) mexicana ssp.] ( Friese, 1899: 151), image: Nemésio, 2009a: 5 a–d, Lectotype 3, designated by Moure, 1967b: 407, ZMHB: NICARAGUA.

Eufriesea convexa [ Euglossa (Eulema!) mexicana ssp.] ( Friese, 1899: 151), Holotype Ƥ, ZMHB: PERU, Loreto, Pebas (not BRAZIL, Tebas).

Eufriesea corusca [ Euplusia ] ( Kimsey, 1977: 10–12, figs. 12, 24, 36), Holotype 3, UCDC: PANAMA, Colón, Barro Colorado Island, leg. Dressler.

Eufriesea dentilabris [ Euglossa (Eulema!)] ( Mocsáry, 1897: 443–444), image: Nemésio, 2009a: 46 a–d, Holotype 3, HNHM: BRAZIL, Espírito Santo.

Eufriesea distinguenda [ Euglossa (Eulema!)] ( Gribodo, 1882: 267–268), image: Nemésio, 2009a: 24 a–e, Holotype Ƥ, MSNG: FRENCH GUIANA, Cayenne.

Eufriesea dressleri [ Euplusia ] ( Kimsey, 1977: 12, figs. 3, 13, 25, 37), Holotype 3, UCDC: PANAMA, Colón, Piña, leg. Dressler.

Eufriesea duckei [ Euglossa (Eulaema) ] ( Friese, 1923: 28), image: Nemésio, 2009a: 6 a–d, Holotype Ƥ, AMNH: BRAZIL, Amapá, Macapá, leg. Ducke.

Eufriesea eburneocincta [ Euplusia ] ( Kimsey, 1977: 12–13, figs. 4, 14, 26, 38), Holotype 3,? (not in USNM, UCDC, UF; paratypes in UCDC, UF): GUYANA, Pomeroon–Supenaam, Dawa, leg. Hamer.

Eufriesea elegans [ Eulaema ] (Lepeletier de Saint Fargeau, 1841: 13), Lectotype Ƥ, designated by Moure, 1967b: 407 (cf. ICZN Art. 74.6), MRSN: FRENCH GUIANA, Cayenne.

Eufriesea excellens [ Euglossa (Eumorpha) ] ( Friese, 1925: 27), Lectotype 3, designated by Moure, 1967b: 408 (cf. ICZN Art. 74.6), ZMHB: ECUADOR, Guayas, Guayaquil, leg. Buchwald.

Eufriesea fallax [ Euglossa ] ( Smith, 1854: 381), Lectotype 3, designated by Moure, 1967b: 408, BMNH 17b1175: BRAZIL, Pará.

Eufriesea flaviventris [ Euglossa (Eulema!)] ( Friese, 1899: 152), Lectotype Ƥ, designated by Moure, 1967b: 408, HNHM: BRAZIL.

Eufriesea formosa [ Euglossa (Eumorpha) ] ( Mocsáry, 1908: 581), Lectotype Ƥ, designated by Moure, 1967b: 408 (cf. ICZN Art. 74.6), HNHM: BRAZIL, Amazonas, Tefé.

Eufriesea fragrocara [ Euplusia ] ( Kimsey, 1977: 13, figs. 5, 15, 27, 39), Holotype 3, UCDC: PERU, Huánuco, Llullapichis, Río Pachitea, leg. Dressler.

Eufriesea fuscatra [ Euplusia danielis ssp.] ( Moure, 1999: 94), Holotype 3, DZUP: BRAZIL, Tocantins, Santa Isabel do Morro, Ilha do Bananal, leg. Alvarenga: Locality originally in the state of Goiás.

Eufriesea heideri Nemésio & Bembé, 2008: 243 –244, figs. 1a, 1c, 1e, 2, Holotype 3, UFMG: BOLIVIA, Cochabamba, Chapare, Chimoré river, close to Entre Ríos, 310 m, leg. Heider.

Eufriesea kimimari González & Gaiani, 1989: 150 , fig. 2, Holotype 3, MIZA: VENEZUELA, Táchira, Río Frio, leg. Romero.

Eufriesea laniventris [ Euglossa ] ( Ducke, 1902a: 403), Lectotype 3, designated by Moure, 1967b: 408, MPEG: BRAZIL, Pará, Belém, leg. Ducke: The nomen appeared first both in a key and accompanied by a brief description (October 1st), thus giving this paper priority over the actual description, Ducke, 1902b: 567, 575 (December).

Eufriesea limbata [ Euglossa (Eulema!)] ( Mocsáry, 1897: 442), Holotype Ƥ, HNHM: BRAZIL, Piauí. basalis [Eulema!] ( Friese, 1898: 203–204), Lectotype Ƥ, designated by Moure, 1967b: 407, ZMHB: SURI- NAME, leg. Fruhstorfer.

Eufriesea longipennis [ Euglossa (Eumorpha) ] ( Friese, 1925: 28), Lectotype Ƥ, designated by Moure, 1967b: 408 (cf. ICZN Art. 74.6), ZMHB: ECUADOR, Guayas, Guayaquil, leg. Buchwald.

Eufriesea lucida [ Euplusia ] ( Kimsey, 1977: 13–15, figs. 16, 28, 40), Holotype 3, UCDC: COLOMBIA, Valle del Cauca, Valle Anchicaya, leg. Kennedy.

Eufriesea lucifera Kimsey, 1977: 18 , figs. 17, 29, 41, Holotype 3, UCDC: PANAMA, Panama, El Llano–Cartí Road, 19 km N of El Llano, leg. Siri.

Eufriesea macroglossa [ Euplusia ] ( Moure, 1965: 274–275), Holotype 3, SEMC: COSTA RICA, Cartago, Turrialba, leg. Dodson.

Eufriesea magrettii [ Euglossa (Eumorpha) ] ( Friese, 1899: 148), Lectotype 3, designated by Moure, 1967b: 408, MSNG: VENEZUELA.

fulvohirta [ Euglossa (Eumorpha) magrettii ssp.] ( Friese, 1899: 148), Lectotype 3, designated by Moure, 1967b: 408, MSNG: VENEZUELA.

Eufriesea mariana [ Euglossa Mariana ] ( Mocsáry, 1896: 4–5, pl. 1, fig. 4), Holotype Ƥ, HNHM: BOLIVIA, El Beni, Bueyes.

superba [ Euglossa ] ( Mocsáry, 1898: 498), Holotype 3, HNHM: BOLIVIA, La Paz, Cordillera Real, Zongo (“Songo”): Junior homonym of superba Hoffmannsegg, 1817 .

tucumana [ Euglossa (Eumorpha) mariana ssp.] ( Schrottky, 1902: 117), Holotype Ƥ, MACN: ARGENTINA, Tucumán.

pulcherrima [ Euglossa (Eufriesea) ] ( Friese, 1923: 27), nom. nov. for Euglossa superba Mocsáry 1898 , nec Hoffmannsegg 1817.

Eufriesea melissiflora [Paleoeuglossa] ( Poinar, 1998: 31–34, figs. 1–5), Holotype Ƥ, WACA: DOMINICAN REPUBLIC (amber inclusion). According to D.W. Roubik, not an Eufriesea due to the presence of scutellar tuft.

Eufriesea mexicana [ Euglossa (Eulema!) Mexicana ] ( Mocsáry, 1897: 444–445), Lectotype Ƥ, designated by Moure, 1967b: 408, HNHM: MEXICO, Veracruz, Presidio.

Eufriesea micheneri Ayala & Engel, 2008: 228 –234, figs. 1–9, Holotype 3, UNAM: MEXICO, Jalisco, Mascota, leg. Ayala.

Eufriesea mussitans [ Apis ] ( Fabricius, 1787: 301), image: Nemésio, 2009a: 50 a–d, Lectotype Ƥ, designated by Moure, 1967b: 408 (cf. ICZN Art. 74.6), ZMUC: FRENCH GUIANA, Cayenne.

inermis [ Euglossa (Eulema!) mexicana ssp.] ( Friese, 1899: 151), Lectotype 3, designated by Moure, 1967b: 408, AMNH (type status unconfirmed): VENEZUELA.

Eufriesea nigrescens [ Euglossa (Eufriesea) magretti ssp.] ( Friese, 1923: 27), nom. nov. for Euglossa nigrita Friese 1899 , nec Lepeletier de Saint Fargeau 1841.

nigrita [ Euglossa (Eumorpha) magrettii ssp.] ( Friese, 1903: 574), Lectotype Ƥ, designated by Moure, 1967b: 409, AMNH (type status unconfirmed): ECUADOR, Bolívar, Balsapamba (Balzapamba), leg. Schulz (durch A Schulz).

Eufriesea nigrohirta [ Euglossa (Eumorpha) ] ( Friese, 1899: 142–143), Nemésio, 2005: 41, 3; image: Nemésio, 2009a: 52a–d, 53d, Holotype Ƥ, ZMHB: BRAZIL, Pará.

faceta [ Euplusia ] ( Moure, 1999: 98–99), image: Nemésio, 2009a: 53a, 54a–d, Holotype Ƥ, DZUP: BRAZIL, Bahia, Mucugê, 984m, leg. Benton.

Eufriesea opulenta [ Euglossa (Eumorpha) ] ( Mocsáry, 1908: 582), Lectotype 3, designated by Moure, 1967b: 409 (cf. ICZN Art. 74.6), HNHM: PERU, Junín, Chanchamayo.

Eufriesea ornata [Eulema!] ( Mocsáry, 1896: 3–4), image: Nemésio, 2009a: 38, Holotype 3, HNHM: BRAZIL, Pará: Purchased through Dr. O. Staudinger’s insect dealership, Dresden, Germany.

Eufriesea pallida [ Euplusia ] ( Kimsey, 1977: 15, figs. 6, 8, 18, 30, 42), Holotype 3, UCDC: MEXICO, Chiapas, Tuxtla Gutiérrez, leg. Dodson.

Eufriesea pretiosa [ Euglossa (Eulema!)] ( Friese, 1903: 575), Syntype Ƥ, Moure, 1967b: 408 (cf. ICZN Art. 74.6),?; “AMNH” (false type): COLOMBIA, Valle del Cauca, Cauca–Tal, near Popayán: Friese cited Ecuador and Caucatal as the type locality. However, Cauca–Tal is in Colombia. The presumably false type from AMNH is from Ecuador (Tungurahua, Santa Ines).

Eufriesea pulchra [ Euglossa ] ( Smith, 1854: 381), image: Nemésio, 2009a: 7 a–d, Holotype Ƥ, BMNH 17b948: BRAZIL, Pará, Tapajós.

Eufriesea purpurata [ Euglossa ] ( Mocsáry, 1896: 5, fig. 5), Lectotype Ƥ, designated by Moure, 1967b: 409, HNHM: PERU, Loreto, Iquitos.

Eufriesea pyrrhopyga Faria & Melo, 2011: 36 –37, figs. 1–5, Holotype Ƥ, DZUP: BRAZIL, Pernambuco, Recife, leg. Lopes.

Eufriesea rufocauda [ Euplusia ] ( Kimsey, 1977: 15–16, figs. 7, 19, 31, 43), Holotype 3, UCDC: PANAMA, Panama, Cerro Campana, leg. Kimsey.

Eufriesea rugosa [ Euglossa (Eumorpha) ] ( Friese, 1899: 152–153), Lectotype Ƥ, designated by Moure, 1967b: 409, ZMHB: MEXICO.

Eufriesea schmidtiana [ Euglossa (Eulema!)] ( Friese, 1925: 26), Lectotype 3, designated by Moure, 1967b: 409, ZMHB: COSTA RICA, Heredia, Puerto Viejo de Sarapiquí, leg. Schmidt (Albert).

Eufriesea smaragdina [ Centris ] ( Perty, 1833: 150, pl. 28, fig. 13), image: Nemésio, 2009a: 30 (org. plate), 33 (org. plate), 55a–d, 57, Holotype 3, ZSM: BRAZIL, Minas Gerais, leg. Spix et al.

Eufriesea superba [ Plusia ] ( Hoffmannsegg, 1817: 53), Holotype 3,?: BRAZIL, Pará. manni [ Eulaema ] ( Cockerell, 1912: 41–42), Holotype 3, USNM 58113: BRAZIL, Rondônia, Abunã, leg. Mann & Baker.

Eufriesea surinamensis [ Apis ] (Linnaeus, 1758: 579 (not 575)), image: Nemésio, 2009a: 28 b, d, 29b, d, f, 60a–f, Holotype Ƥ, NHRS: SURINAME, leg. Rolander: (not Linnaeus, 1758: 575). See Linnaeus, 1767: 961 for First Reviser action and van der Vecht, 1959: 211 for a discussion of the priority of this nomen. tropica [ Apis ] ( Linnaeus, 1758: 579), image: Nemésio, 2009a: 28 a, 29a, c, e, Holotype Ƥ, UUZM: “Calidis regionibus” (possibly SURINAME).

abdomenflavum [ Apis abdomen flavum] ( De Geer, 1773: 574, pl. 28, figs. 9, 10), image: Nemésio, 2009a: 34 (org. plate), Holotype Ƥ, NHRS: SURINAME: Possibly a syntype.

angulata [ Euglossa ] ( Mocsáry, 1897: 443), Lectotype Ƥ, designated by Moure, 1967b: 406, HNHM: BRAZIL, Piauí.

bruesi [ Eulaema ] ( Cockerell, 1914: 307–308), image: Nemésio, 2009a: 30c, Holotype Ƥ, USNM 58114: ECUADOR, Guayas, Guayaquil, leg. Brues.

amabilis [ Eulaema ] ( Cockerell, 1917a: 476–477), image: Nemésio, 2009a: 30b, 59, Holotype 3, USNM 23155: BRAZIL, Amazonas, Manaus, leg. Merrill.

tectora [ Euplusia ] ( Kimsey, 1977: 16, figs. 20, 32, 44), Holotype 3, UCDC: PANAMA, Colón, Frijoles, leg.

Dressler.

Eufriesea theresiae [ Euglossa (Eumorpha) Theresiae ] ( Mocsáry, 1908: 581–582), image: Nemésio, 2011a: 1, 2, Lectotype Ƥ, designated by Moure, 1967b: 409–410 (cf. ICZN Art. 74.6), HNHM: BRAZIL, Amazonas, Tefé.

Eufriesea velutina [ Euplusia ] ( Moure, 1999: 101–103), Holotype Ƥ, DZUP: VENEZUELA, Mérida.

Eufriesea venezolana [ Centris ] ( Schrottky, 1913: 708), Syntype 3, MPSP: VENEZUELA.

Eufriesea venusta [ Euplusia ] ( Moure, 1965: 273–274), Holotype Ƥ, SEMC: PANAMA, Panama, Cerro Campana, leg. Hanson.

Eufriesea vidua [ Euplusia ] ( Moure, 1976: 277–281), Holotype Ƥ, HNHM: SURINAME. xantha [ Euplusia ] ( Kimsey, 1977: 17–18, figs. 21, 33, 45), Holotype 3, USNM: GUYANA, Upper Demerara – Berbice, Mazaruni–Potaro river, Rockstone, leg. Williams.

Eufriesea violacea [ Euglossa ] ( Blanchard, 1840: 405, pl. 7, fig. 3), image: Nemésio, 2009a: 61 (org. plate), 63a–d, Neotype 3, Nemésio, 2009a: 69, fig. 63, UFMG: BRAZIL, Minas Gerais, Viçosa, leg. Soares.

Eufriesea violascens [ Euglossa ] ( Mocsáry, 1898: 497), image: Nemésio, 2009a: 15 a–f, Lectotype Ƥ, designated by Moure, 1967b: 410, HNHM: BOLIVIA, La Paz, Cordillera Real, Zongo (“Songo”).