Ochthephilus forticornis (Hochhuth, 1860)

Makranczy, György, 2014, Revision of the genus Ochthephilus Mulsant & Rey, 1856 (Coleoptera: Staphylinidae, Oxytelinae), Revue suisse de Zoologie 121 (4), pp. 457-694 : 669-672

Ochthephilus forticornis (Hochhuth, 1860)

NON-TYPEMATERIALEXAMINED: NORWAY: Kåfjord, Lyngen *+69.92/+20.20*, (VI), exp. Münster, ZMUN (5). – RUSSIA: Kamtschatka, Bolscherjetsk [Kavalerskoye] *+52.92/+156.58*, (VII), MZHF (1). – Khabarowskiykray, Badshalskiyrange, endofriver Omot-Makit, taiga, 850m *+50.58/+134.32*, (VII), coll. Schülke (1). – PrimorskijKraj, Lazovskij Distr., river Elomovsky [= Primorye Terr., S Sikhote-Alin ridge, Partizansky ridge, Lysajamt., Elamovskyriver], 650m *+43.26/+133.70*, (VII), coll. Pütz (34). – SouthPrimorye, Lazovskijdistrict, Lazo *+43.38/+133.90*, (VII), coll. Shavrin (2). – Primorie, Schutzgebiet Lazowskij, Kl. S-j. Log *+43.36/+133.87*, (VI), coll. Schülke (1). – Приморскийкрай, Хасанскийрайон, заповедникКедроваяПадь, *+43.14/+131.61*, (VII-VIII), ZMUM (2). – Ussuri-Gebiet, RajonPartizansk, Poworotnaja-Tal *+42.68/+133.03*, (VI), ZMHB (42). – Primorsk, Ussuriysk, Kaimanovka [Каймановка], 150-300m *+43.63/+132.22*, (VII), ZMUM (1). – PrimorskiyKray, Sikhote-Alin, Biol. Stat., 30kmSEChuguyevka, 44°05'N, 134°12'E, 650m, (V-VI), SDEI (25). – QuellgebietdesIrkut *+52.12/+100.87*, coll. Kraatz, SDEI (2), in part coll. Skalitzky and coll. Eppelsheim, NHMW (7). – Бурятская АССР, курорт Аршан, *+51.95/+102.50*, (VII-VIII), coll. Shavrin (3). – N-Cisbaikalia, Buryatia, BaikalMts., Kurkula [Куркула] river, 55°05'52.0"N, 108°55'33.9"E, 725m, (VII), coll. Shavrin (2). – Khamar-Daban Mts., KomarinskiyMts., middleflowofSlyudyankaRiv., 51.587736°N, 103.630056°E, (VII), coll. Shavrin (1). – хребетХамар-Дабан, верховьярекиОсиновки, 1500 м *+51.50/+104.71*, (VII), coll. Shavrin (1). – Красноярский край, Саян-Шушенский заповедн., р. Большие Уры близ устья р. Оттуг-Суук [Krasnoyarsk Territory, Yermakovskiy District, Sayano-Shushenskiy Biosphere Reserve, Bolshiye Ury River basin near Ottug-Suuk (= Otuk-Sug) river mouth, 51°59'N, 91°38'E] (V), HNHM (1). – Transbaikalia, ChitaArea, KyrenskiyDistrict, Bal'dzhariver, 49°17'54.4"N, 110°09'20.8"E, 1432m, (VIII), coll. Shavrin (2). — CHINA: Gansu, Xinlong Shan, cca. 70kmSLanzhou, 2225-2380m *+35.07/+103.76* (C32), (VII), MHNG (1$), Shanxi, Wutaishan *+39.07/+113.58*, (VI), coll. Rougemont, OXUM (6), Beijing, Xizolongmen *+40.25/+116.50*, (VI), coll. Rougemont, OXUM (1), QinghaiProv., DabanShan, 62kmNW Honggu, creekvalley, 36°51'15-28"N, 102°36'34"-37'07"E, 2236-2350m (CH11-09e), (VII), coll. Schülke (1£). — MONGOLIA: Гоби-Алтайский аймак, 25км ЮВ айм. Алтая (Юсун- Булака), *+46.23/+96.50*, (VII), ZISP (1). – Bayanhongor, 17кмЮШине-Джинста. горы. родник *+44.50/+99.27*, (VI), coll. YanushevviaM. Gildenkov (2). – Bajanchongoraimak, Changaj [Khangay] Gebirge, Somon Zag am fluss Zag [Заг] Gol, 2100m (709) *+47.11/+99.12*, (VII), exp. Kaszab, HNHM (5), coll. Smetana, CNCI (1). – Uvsaimak, am Fluss Baruunturuun gol neben Somon Baruunturuun [Баруунтуруун], 1280m (1012) *+49.65/+94.38*, (VI), exp. Kaszab, HNHM (1). – Bayan-Ölgiyaimak, imTaldesFlusses Chavcalyn gol, 24km O von Somon Cagannur [Tsagaannuur, Цагааннуур], 1890m (1043) *+50.11/+105.81*, (VI), exp. Kaszab, HNHM (2), coll. Smetana, CNCI (1). — SOUTH KOREA: Jeju-do, Seogwipo-si *+33.25/+126.56* (II), CNUK (1). — JAPAN: Honshu, Gunma pref., KonseiPass, ca. 1800-1900m *+36.81/+139.35*, (VIII), BMNH (1). – Honshu, Gunma pref., Mt. Hotaka (foot), ca. 1300m *+36.78/+139.16*, (VIII), BMNH (13, 1♀). – Iwatepref., Kawara-bo, Mt. Hayachine *+39.45/+141.42*, (VI), coll. Naomi, KUEC (3). – Tokushimapref., Mt. Tsurugi *+33.85/+134.10*, (VI), coll. Naomi, KUEC (1). – Tottoripref., Mt. Daisen *+35.37/+133.54*, (VI), coll. Naomi, KUEC (3). – Gifupref., Hirayu *+36.15/+137.50*, (VI), coll. Naomi, KUEC (1). – Tochigipref., Nikko, NikkoYumoto *+36.81/+139.41*, (VI), coll. Naomi, KUEC (1). – Gunmapref., Marunuma *+36.80/+139.36*, (VII), coll. Naomi, KUEC (43). – Yamanashipref., Jigokudani, Mt. Yatsu *+35.88/+138.51*, (VII), coll. Naomi, KUEC (3). – Yamanashipref., Mt. Daibosatsu, Kaminikkawapass *+35.72/+139.00*, (VII), coll. Naomi, KUEC (31). – Yamanashipref., Hirogawara, Mt. Shirane *+35.66/+138.30*, (VII), coll. Naomi, KUEC (16). – Narapref., Ohdaigahara *+33.91/+135.86*, (VI), coll. Naomi, KUEC (7). – Ehimepref., Mt. Odami [Odamiyama] *+33.54/+132.85*, (VI), coll. Naomi, KUEC (4). – Hokkaido, Mt. Hakken-zan *+42.96/+141.31*, (VI), coll. Naomi, KUEC (1). – Kyotopref., Ashiu [Natural Forest] *+35.31/+135.72*, (VI), coll. Ito (3). – Nara pref., Mt. Ohdai *+33.91/+135.86*, (VandVI), coll. Ito (4). – Narapref., Dorogawa *+34.27/+135.88*, (V), coll. Ito (3). — USA: OREGON: BakerCo., PineCreeknearBaker, BlueMountains *+44.82/- 118.08*, (VI), FMNH (34). – Grant Co., E of Prairie City, Blue Mountains, E of Dixie Pass, LunchCreek (2 miW), c. 5000ft *+44.58/-118.44*, (V-VI), FMNH (3). – CurryCo., Humbug MountainStatePark *+42.69/-124.43*, (VII), FMNH (2). – ClackamasCo., Mt. HoodNational Forest, StillCreekCampground, 3700' *+45.30/-121.77*, (VII), FMNH (1). – IDAHO: Blaine Co., SawtoothNationalForest, PrairieCreek, Hwy. 75 *+43.95/-114.65*, (VII), CNCI (2). – IdahoCo., 20.7miWSWLoloPass, SquawCreek, 3150ft *+46.49/-114.84*, (VII), AMNH (49). – MONTANA: MissoulaCo., 27miWSWLolo, WestForkLoloCreek, 4200ft *+46.71/- 114.53*, (VII), AMNH (10). – MissoulaCo., 1.5miSWLoloHotSprings, LeeCreek, 4200ft *+46.71/-114.54*, (VII), AMNH (5). – CarbonCo., PalisadesCampground, 3miWRedLodge, 5400' *+45.17/-109.31*, (VIII), CNCI (1). – CALIFORNIA: TrinityCo., ShastaNationalForest, DeadfallCreek, 7000' *+41.32/-122.51*, (VII), CNCI (1). – ElDoradoCo., 3miEKyburz, 5500' *+38.78/-120.24*, (VIII), CNCI (1). – SiskiyouCo. 5.4miSESeiadValley, O'NeilCreek, 1500ft *+41.81/-123.12*, (VI), AMNH (16). – AlpineCo. 26.5miNEStrawberry (TuolumneCo.), near ClarkForkRiver, 6400ft *+38.58/-119.69*, (VII), AMNH (5). – TuolumneCo. 33kmNE Starwberry, nearDeadmanCreek, 8900ft *+38.32/-119.70*, (VII), AMNH (1). – NEVADA: LanderCo., 16miSAustin, BigCreekCanyon, 7100ft *+38.15/-115.72*, (VI), AMNH (1). – LanderCo., 17miSAustin, BigCreekCanyon, 7400ft *+38.14/-115.72*, (VI), AMNH (1). – WhitePineCo., 8miWBaker, BakerCreekCamp[ground], 7700ft *+38.99/-114.24*, (VIII), AMNH (1). – UTAH: Washington Co., SE of Pine Valley National Forest, Trail 16, Earl Spring, 7100' *+37.38/-113.55*, (VI), AMNH (34). – GarfieldCo., HenryMts., LonesomeBeaver, 7500' *+38.12/-110.75*, (VII), CNCI (1). – Manti-LaSalNationalForest, WarnerCampground, 9200' *+38.52/-109.28*, (VII), BMNH (7). – COLORADO: DeltaCo., Hwy 65, 11kmNWjnct. For. Rd. 100, GrandMesaN.F., 9400ft *+39.10/-108.15*, (VII), CNCI (1). – EagleCo., 6km NWDotsero, DeepCreek, 6700' *+39.68/-107.09*, (VII), CNCI (2). – MontezumaCo., 28.5mi NEDolores, SchoolhouseDraw, 8000ft *+37.74/-108.16*, (VII), AMNH (9). – MontezumaCo., 25miNEDolores, PriestGulchSpring, 7800ft *+37.58/-108.17*, (VII), AMNH (5). – Chaffee Co., 9miWBuenaVista, CottonwoodCreek, 9600ft *+38.81/-106.27*, (VII), AMNH (1). – San JuanCo., 2-3miSSilverton, 9,900-10,000ft *+37.76/-107.67*, (VIII), AMNH (6). – Huerfano Co., 15miSSWLaVeta, CucharasCreek, 9400ft *+37.35/-105.10*, (VII), AMNH (2). – HuerfanoCo., 8miSSWLaVeta, CucharasRiver, 7800ft *+37.41/-105.06*, (VIII), AMNH (1). – OurayCo., BoxCanyonFalls, Ouray, 7800ft *+38.03/-107.66*, (VII), AMNH (13), coll. Bernhauer, FMNH (1). – ARIZONA: YavapaiCo., CampVerde, VerdeRiver *+42.28/-124.39*, (IV), AMNH (1). – GrahamCo., PinalenoMts., GrantCreek, 8500' *+32.68/-109.93*, (VII), CNCI (11), MHNG (1). – GrahamCo., PinalenoMts., PostCreek, 9000' *+33.04/-110.36*, (VII), CNCI (11). – ChuskaMts., ApacheCo., WagonwheelCampground, 2250m *+36.05/- 109.18*, (VII), CNCI (34), BMNH (30). – ChuskaMts., ApacheNationalForest, ApacheCo., EscudillaMt., 2840m, 9miNEAlpine *+33.92/-109.11*, (VII), CNCI (8). – CochiseCo., ChiricahuaMtns., RuckerCanyon, (1730-) 1760m *+31.75/-109.35*, (VII), CNCI (1), BMNH (1). – GreenleeCo., HannaganCamp *+33.64/-109.33*, (VII), CNCI (1). – NEWMEXICO: OteroCo., LincolnNationalForest, 13miSCloudcroft, 7500' *+32.80/-105.74*, (VII), CNCI (142), MHNG (7). – OteroCo., LincolnNationalForest, 1miSECloudcroft, 8750' *+32.94/- 105.72*, (VII), CNCI (3). – LincolnCo., LincolnNationalForest, SierraBlancaSkiArea, 10600' *+33.40/-105.79*, (VII), CNCI (9), MHNG (1). – LincolnCo., SierraBlanca, 9500' *+33.37/-105.80*, (VII), CNCI (3). – LincolnCo., S. ForkBonitoCreek, ca 7miWAngus, 7500-8000' *+33.44/-105.76*, (VII), FMNH (2). – SandiaMts., BernalilloCo., CibolaNational Forest, LasHuertasCreek, 7000' *+35.23/-106.41*, (VII), CNCI (18). – SandiaMts., Bernalillo Co., CibolaNationalForest, LasHuertasCreek, 6500' *+35.24/-106.41*, (VII), CNCI (4). – SandiaMts., BernalilloCo., CibolaNationalForest, TreeSpringTrail, 8500' *+35.18/-106.42*, (VII), CNCI (9). – [El] Porvenir *+35.71/-105.41*, coll. Bernhauer, FMNH (1). – SOUTHDA- KOTA: LawrenceCo., 13miSSWSpearfish, RoughlockFalls, 5600ft *+44.35/-103.94*, (VII), AMNH (5). – LawrenceCo., 16miSSWSpearfish, LittleSpearfishCreek, 6000ft *+44.29/- 104.00*, (VII), AMNH (42). – LawrenceCo., 5miSofSpearfish, BridalVeilFalls, 4700ft *+44.42/-103.87*, (VII), AMNH (1). – MISSOURI: RipleyCo., 5.5miNBriar *+36.72/-90.97*, (V), AMNH (2). – INDIANA: 1miNNWofDeputy, Foster'sCaves *+38.81/-85.66*, (VI), AMNH (3). – KENTUCKY: AdairCo., 3miWNWColumbia, Todd'sCave *+37.11/-85.37*, (VI), AMNH (1). – NEWYORK: LakePlacid *+44.28/-73.98*, (VII), CASC (1). – ALASKA: Fairbanks *+64.85/-147.72*, (VII), USNM (1). – KenaiMts., TernLakeCampground, 700' *+60.53/-149.54*, (VI), CNCI (2). – KenaiPen., AnchorRiver, campground 12miNHomer, 450' *+59.77/-151.50*, (VI), CNCI (2). – mi24 WalesHwy., HessCreek, 600' *+65.67/- 149.17*, (VII), CNCI (1). – GeorgeParksHwy, mi220 *+63.83/-149.05*, (VII), CNCI (9). – KenaiPeninsula, 9miNSeward *+60.23/-149.42*, (VII), AMNH (1). — CANADA: YUKON TERRITORY: DempsterHwy., mi154, OgilvieRiver, 1900' *+65.70/-138.07*, (VII), CNCI (3). – DempsterHwy., mi100.5, EngineerCreek, 3000' *+65.36/-138.20*, (VII), CNCI (5), HNHM (2). – DempsterHwy., mi139.5, 1900' *+65.58/-138.17*, (VII), CNCI (40), HNHM (1). – DempsterHwy., mi45, 3500' *+64.48/-138.23*, (VII), CNCI (42). – BritishMts., "TealLake", at edge of Firth R. (84-4C), 68°54'N, 140°23'W, 400m, (VI), CNCI (2). – BRITISH COLUMBIA: 'KsanHistoricVillagenearHazelton, 55°14.95'N, 127°40.64'W, 218m (VII), CNCI (1). – Ymir *+49.29/-117.21*, (VI), CNCI (1). – AlaskaHwy., mi445, PetersonCreek *+59.00/-125.78*, (VII), CNCI (29). – AltaLake, 2200' *+49.23/-125.65*, (VIII), CNCI (1). – 21miWCreston *+49.10/-116.95*, (VI), CNCI (5). – 8miWCreston *+49.10/-116.68*, (VI), CNCI (16). – 7miETerrace *+54.52/-128.41*, (VI), CNCI (2). – SlideMt., 10miEBarkerville 4500' *+52.67/-121.87*, (VI), CNCI (6). – Fernie, LizardCreek *+49.47/-115.08*, (VII), CASC (1) [a misidentified paratype of Ancyrophorus columbiensis ]. – Two Sisters Mt., 15mi E Barkerville, 4500' *+53.20/-121.53*, (VI), CNCI (12). – ManningProvincialPark, 20miEHope *+49.38/-120.87*, (VI), CNCI (21), HNHM (1). – ManningProvincialPark, 5kmEwestPark entrance *+49.24/-121.13*, (VI), CNCI (2). – ManningProvincialPark, DerekFalls 4000' *+49.07/-120.93*, (VII), CNCI (1). – ManningProvincialPark, BlackwellPeak, 6000-6750' *+49.18/-120.82*, (VI), CNCI (1). – YohoN. P., valleyofHagenPeak *+51.58/-116.72*, (VIII), CNCI (3). – YohoN. P., AmiskwiRiver, 6000' *+51.38/-116.55*, (VIII), CNCI (10). – Otterhead R., YohoN. P., 5500' *+51.41/-116.69*, (VIII), CNCI (2). – KootenayN. P., SinclairCreek, 5000' *+50.64/-116.09*, (VI), CNCI (53). – KootenayN. P., KimptonCreek, 4000' *+50.65/- 115.97*, (VII), CNCI (2). – Mt. RevelstokeN. P., 2000' *+51.05/-118.13*, (VIII), CNCI (1). – GlacierN. P., 10 miERogersPass *+51.30/-117.32*, (VI), CNCI (2). – 17kmNWhistler (Hwy99), 600m (35a) *+50.28/-122.93*, (VII), MHNG (1). – ALBERTA: JasperNationalPark, MietteHotSprings, SulphurCreek, 4500ft *+53.15/-117.78*, (VII), AMNH (4). – Jasper, WhistlerCreek *+52.85/-118.09*, coll. Cameron, BMNH (5). – SprayLakes *+50.90/-115.34*, (VII), coll. Cameron, BMNH (1). – L. Minnewanka *+51.28/-115.40*, (VII), coll. Cameron, BMNH (17). – Edmonton *+53.53/-113.50*, (IX), coll. J.W. Green, CASC (1). – Banff, (Sundance) *+51.17/-115.62*, (VI), coll. Bernhauer, FMNH (2), coll. Cameron, BMNH (10). – 11miSWBanff, HealyCreek *+51.10/-115.81*, (VIII), CNCI (17). – BanffN. P., MoraineLake, 6200' *+51.32/-116.18*, (VI), CNCI (2). – BanffN. P., CirrusMt., 5500' *+52.17/-116.99*, (VI), CNCI (1). – BanffN. P., BoomLake, 6000' *+51.27/-116.08*, (VII), CNCI (1). – BanffN. P., Mt. Temple, skilodge, 6600' *+51.35/-116.22*, (VIII), CNCI (16). – 1miEKananaskisF. E. S., LuskCreek, *+51.03/-115.03*, (VIIandVIII), CNCI (22). – 1miEKananaskisF. E. S., 6000' *+51.03/-115.03*, (VII), CNCI (2). – MarmotCreekBasin, 10miSWKananaskisF. E. S., 6500' *+50.95/-115.13*, (VII), CNCI (1). – WatertonLakesN. P., RoweCreek, 5400ft *+49.06/- 114.02*, (VIII), CNCI (22). – WatertonLakesN. P., RoweCreek, 5460ft *+49.06/-114.02*, (VIII), CNCI (12), BMNH (44). – WatertonLakesN. P., CryptLake, 6500' *+49.00/-113.84*, (VI), CNCI (2). – WatertonLakesN. P., mi3 ChiefMt. Hwy, 4500' *+49.09/-113.79*, (VI), CNCI (1). – WatertonLakesN. P., CameronCreek, 4900' *+49.04/-114.00*, (VI), CNCI (8). – WatertonLakesN. P., CameronCreek, 5200' *+49.03/-114.01*, (VI), CNCI (9). – Waterton LakesN. P., RoweLakesTrack, 6300-6400' *+49.05/-114.05*, (VI), CNCI (9). – Waterton LakesN. P., LowerBerthaFalls, 4800' *+49.04/-113.90*, (VI), CNCI (5). – WatertonLakesN. P., RoweCreekTrack, 5300' *+49.06/-113.99*, (VIII), CNCI (3). – ONTARIO: SouthMarchnr. Ottawa (256) *+45.38/-75.90*, (V), MHNG (13, 2). – NEWBRUNSWICK: Northumberland Co., N.W. MiramichiRiver *+46.95/-65.75*, (VII), CNCI (13).

Ochthephilus lenkoranus ( Scheerpeltz, 1950) NON-TYPEMATERIALEXAMINED : SWITZERLAND: Kt. Vaud, lesAvants *+46.45/

+06.94*, (VII), MHNG (1). — ITALY: Crognaleto ( TE), Zingano *+42.59/+13.49*, ( VIII), coll. Zanetti (1) . – Cortino ( TE), Padula *+42.62/+13.47*, (X), coll. Zanetti (1) . – Sicilia, NW Francavilla , 37°56'N, 15°06'E, 480m, (V), coll. Assing (1) . — SLOVAKIA: Pieniny, Dunajec river *+49.39/+20.41*, coll. Roubal , SNMC (1). – NováSedlica, Zbojskýstream 450m *+49.04/+22.48*, ( VI), SMBC (13) , HNHM (13). – MaláFatra, Vratnadolina, Vrátnanka *+49.25/+19.04*, ( VIII), ZMHB (1). – Bukovskévrchy, bankofZbojskýstream, 450m *+49.04/+22.47*, ( VI), SMBC (13, 4) , HNHM (23, 1♀, 1). – Bukovskévrchy, Ruské , bankof stream, 500m *+49.04/+22.41*, ( VII), SMBC (2$) , HNHM (13, 1♀). – Bardejov, Rhodyhes 3 dvor *+49.29/+21.28*, (V), SMBC (1) . — ROMANIA: jud. Caraş-Severin, MunţiiSemenic, CheileMinişului, 4kmSEAnina, p. Miniş , gravelislands (341), 45°01' 48"N, 21°51' 41"E, 620m, (X) , HNHM (1). — BOSNIA-HERZEGOVINA: Hercegovina, Gacko *+43.17/+18.53*, ( VIII), coll. Lohse , MHNG (4). – [Hercegovina, Prenjplanina] Igbar [Idbar] Thal [Grabovicaschlucht] *+43.64/+17.88*, coll.? Zoufal , PMSL (13). – Bosnia, Zvijezda Planina *+44.18/+18.41*, ( VII), coll. Fodor , HNHM (1). – Bosnia, Prozor *+43.82/+17.61*, coll. Leonhard , NHMW (5). — BULGARIA: Melnik *+41.52/+23.40*, (V) , SMNS (1). — ALBANIA: Nord-Albanien, Pukë-FushëiArrësit *+42.05/+19.90*, (V), coll. Smetana , CNCI (3). – Nord-Albanien, FushëiArrësit *+42.06/+20.02*, (V), coll. Smetana , CNCI (1). – Alban. mer., Terbaci [Tërbaçi] *+40.25/+19.65*, (V) , MHNG (3). — GREECE: Vikonsgorge *+39.96/+20.70*, (V), coll. Mahler (3) . – Fokis, 25kmSWLamía, Stromi , riverbank (13), 38°41'24"N, 22°12'54"E, 640m, ( IV), coll. Wunderle (1) . – Epirus, NomósIoánnina, Tímfi Gebirge, beiPapingó, 900m, Laubwald *+39.97/+20.68*, (V) , SMNS (3). — TURKEY: Antalya, Güneycik, bankofKarpuzriver, 600m *+36.75/+31.77*, (I), coll. Assing (2) . – Antalya, Yaylaalan , 900m *+36.95/+31.48*, ( XII), coll. Assing (7) , HNHM (3). – Antalya, umg. Manavgat, 900m [ Yaylaalan ] *+36.93/+31.47*, ( XII), coll. Assing (6) . – umg. Manavgat, Yaylaalan , 850m *+36.95/+31.48*, ( XII), coll. Wunderle (2) . – Antalya, Köprülü-Kanyon, nahe Beskonak , UferdesKöprüÇayı, 126m (008) *+37.11/+31.19*, ( III), coll. Rose (1) . – Antalya, 5kmNWKemer, felsigesFlussuferinSchluchttal (030a), 36°36'03.0"N, 30°29'03.6"E, 197m, ( III) , HNHM (1). – Antalya, Köprülü-Kanyon , 10kmNBeskonak (050a), 37°13'10.9"N, 31°12'45.1"E, 265m, ( III), coll. Rose (26) , SEMC (8), Antalya, Köprülü-Kanyon, 1kmN Beskonak (051), 37°11'14.4"N, 31°10'47.3"E, 178m, ( III), coll. Rose (13) , SEMC (3). – Güneycak , nearGündogmus, 700m *+36.80/+32.02*, (I), coll. Wunderle (1) . – Kastamonu prov., EInebolu (9) *+41.97/+33.77* (V), NHMW (13). – Sinopprov., WYenikonak (11) *+41.87/+34.61* (V) , NHMW (1♀). – Artvinprov. , WBorçka (45) *+41.36/+41.67* ( VI) , NHMW (13). – Erzincanprov. , SERefahiye (77) *+39.89/+38.77* ( VI) , NHMW (13). — GEORGIA: Zchneti [ Tsqneti ] pr. Tbilisi, 800m *+41.71/+44.70*, ( VI), coll. Schülke (23) , HNHM (1♀). — AZERBAIJAN: Lenkoran *+38.78/+48.60*, coll. Reitter , HNHM (2). – NКуткашена, бер. р. Гамзаличай, 1100-1200м *+41.03/+47.85*, (VII), coll. Solodovnikov, ZMUC (1). — RUSSIA: Крым [= Крымская область], Бахчисарайский район, Счастливое, *+44.57/+34.07*, (V), coll. Gusarov, ZMUN (1). – Crimea , CrimeanGamePreserve, upperpart ofKatcha [Кача] river, bank, 800m [44°43'31"N, 33°32'55"E], coll. Schülke (2).