Digitonthophagus bonasus

Génier, François & Moretto, Philippe, 2017, Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae), Zootaxa 4248 (1), pp. 1-110 : 15

Digitonthophagus bonasus species group

Diagnosis. Male cephalic horns long, obliquely divergent in frontal view ( Figs. 24, 26, 28 ); females with horns on vertex ( Figs. 71–73 ).

Figures

FIGURES 23 – 30. 23, 25, 27, 29. Male head, dorsal view; 24, 26, 28, 30. Male head, frontal view. 23 – 24. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 25 – 26. D. uks, holotype; 27 – 28. D. sahelicus, holotype; 29 – 30. D. catta, alloreferent (Tamil Nadu State, India).

FIGURES 71 – 78. Female head, dorsal view. 71. Digitonthophagus bonasus, non type (Tamil Nadu State, India); 72. D. uks, allotype; 73. D. sahelicus, allotype; 74. D. catta, non type (India); 75. D. eucatta, allotype; 76. D. falciger, allotype; 77. D. biflagellatus, allotype; 78. D. gazella, non type (Antisiranana Province, Madagascar).