Digitonthophagus lusinganus

Génier, François & Moretto, Philippe, 2017, Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae), Zootaxa 4248 (1), pp. 1-110 : 57

Digitonthophagus lusinganus species group

Diagnosis. FLP sclerite ventral portion with one or two medioventral carinae ( Fig. 184 A); right lateral fold produced into a rather large everted and open apically conical process with irregular apical edge ( Figs. 165, 168 , 171, 174, 177 ); FLP sclerite subapicoventral lobe acute apically ( Figs. 167 , 176 ); axial and subaxial sclerite rather short, with a distinct transverse uniting sclerotized area medially ( Fig. 171, 174, 177 ).

Figures

FIGURES 183 – 190. Digitonthophagus bonasus, female head, frontal view (Phongsaly Province, Laos); 184. D. aksumensis, FLP sclerite, right lateral view; 185. D. gazella, right lateral fold, ventral view; 186. D. viridicollis, right lateral fold, ventral view; 187. D. eucatta, hypomeron, ventral view, A. anterior hypomeral carina, B. anterior hypomeral depression; D. gazella, hypomeron, ventral view, A. anterior hypomeral carina, B. anterior hypomeral depression; 189. D. namaquensis, vertex dorsal view; 190. D. gazella, vertex dorsal view.

FIGURES 159 – 170. Internal sac sclerites (FLP, A + SA). 159, 162, 165, 168. Dorsal view; 160, 163, 166, 169. Ventral view. 161, 164, 167, 170. Right lateral view. 159 – 161. Digitonthophagus petilus, holotype; 162 – 164. D. viridicollis, holotype; 165 – 167. D. aksumensis, holotype; 168 – 170. D. dilatatus, holotype.

FIGURES 171 – 182. Internal sac sclerites (FLP, A + SA). 171, 174, 177, 180. Dorsal view; 172, 175, 178, 181. Ventral view. 173, 176, 179, 182. Right lateral view. 171 – 173. Digitonthophagus fimator, holotype; 174 – 176. D. lusinganus, lectotype; 177 – 179. D. ulcerosus, holotype; 180 – 182. D. namaquensis, holotype.