Anthocerotaceae

Villarreal, Juan Carlos, Cargill, D. Christine, Söderström, Lars, Hagborg, Anders & Konrat, Matt Von, 2015, Notes on Early Land Plants Today. 70. Nomenclatural notes in hornworts (Anthocerotophyta), Phytotaxa 208 (1), pp. 92-96 : 92-93

Anthocerotaceae

Anthoceros cristatus Steph., Sp Hepat. [Stephani] 5: 996, 1916 ( Stephani 1916). Type:— ASCENSION. Herb. Bescherelle, pores from holotype Stephani herb. no. 24501 (G-00045042! [http://www.ville-ge.ch/musinfo/bd/ cjb/chg/adetail.php?id=141014&lang=en]), lectotype here designated [or possibly holotype, cf. McNeill 2014]) Fig. 1A .

Anthoceros expansus (Steph.) J.C.Villarreal et Cargill , comb. nov. Basionym: — Aspiromitus expansus Steph., Sp. Hepat. [Stephani] 5: 961, 1916 ( Stephani 1916). Type:— CAMEROON. South-West: Buea, 24 September 1891, Preuss 989, Stephani herb. no. 20510 (G-00045055! [http://www.ville-ge.ch/musinfo/bd/cjb/chg/adetail. php?id=141040&lang=en], lectotype here designated [or possibly holotype]).

Anthoceros myriandroecius Steph., Wiss. Ergebn. Deut. Zentr. -Afr. Exped. (1907-08), Bot. 2: 134 ( Stephani 1911 [1910]).Type:— RWANDA.Kiwu lake, Exped. Fr. Mildbraed 808 (G-00128028! [http://www.ville-ge.ch/musinfo/ bd/cjb/chg/adetail.php?id=133358&lang=en], lectotype here designated [or possibly holotype]). Note:—The specimen was used by Stephani for his illustrated icones inedit and it was annotated by Hasegawa in 1993 as “type” but it seems that he did not publish it.

= Anthoceros volkensii Steph., Sp. Hepat. [Stephani] 5: 997, 1916 ( Stephani 1916), syn. nov. Type:— TANZANIA. Kilimanjaro, 17.12.1893, Volkens 1504 (G-00045053! [http://www.ville-ge.ch/musinfo/bd/cjb/chg/adetail. php?id=141027&lang=en], lectotype here designated [or possibly holotype]). Note:—The specimen is annotated by Hasegawa in 1993 as “type” (but it appears that he did not publish it) and conspecific with Anthoceros myriandroecius .

Anthoceros punctatus L., Sp. Pl. 1: 1139 ( Linnaeus 1753). Type:—Herb. Dillen., Oxford, Anthoceros minor , in den oberen Ecken des Blättchens. Hist. musc. (1741) Tab. 68, Fig. 1 . (lectotype designated by Proskauer 1958)

= Aspiromitus dismieri Steph., Sp. Hepat. [Stephani] 5: 961, 1916 ( Stephani 1916), syn. nov. Type:— MADAGASCAR. s. loc. spec., Spores from holotype, Rotereau s.n, Stephanie herb. no.00045054 [http://www.ville-ge.ch/musinfo/ bd/cjb/chg/adetail.php?id=141039&lang=en], lectotype here designated [or possibly holotype]) Fig. 1B . Note:— The type specimen also includes an un-named Phaeoceros species which Hasegawa annotated in 1995.

Anthoceros subtilis Steph., Sp. Hepat. [Stephani] 5: 1003, 1916 ( Stephani 1916). Type:— INDIA. Karnataka: Mangalore, R. Pfleiderer 12923, Stephani herb. no. 12823 (G-00060882 [http://www.ville-ge.ch/musinfo/bd/cjb/ chg/adetail.php?id=115887&lang=en] lectotype designated by Asthana & Srivastava 1991: 54).

= Anthoceros chevalieri Steph., Spec. Hepat. [Stephani] 6: 425, 1923 ( Stephani 1923), syn. nov. Type:— CENTRAL AFRICAN REPUBLIC. Ubangi, 1902, Chevalier 43 (G-00128024! [http://www.ville-ge.ch/musinfo/bd/cjb/chg/ adetail.php?id=133363&lang=en], lectotype here designated [or possibly holotype]).

Folioceros argillaceus (Steph.) J.C.Villarreal et Cargill , comb. nov. Basionym:— Aspiromitus argillaceus Steph., Sp. Hepat. [Stephani] 5: 970, 1916 ( Stephani 1916). Type:— INDONESIA. Java: Bogor (“Buitenzorg”), February 1894, Schiffner s.n., Stephani herb. no. 20494 (G-00061272, lectotype here designated; G-00061271 [http://www. ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=130334&lang=en], G-0061270, JE-04000311, isolectotypes). ≡ Anthoceros argillaceus (Steph.) Verd., Ann. Bryol. 5: 143, 1932 ( Verdoorn 1932).

Folioceros dilatatus (Steph.) J.C.Villarreal et Cargill , comb. nov. Basionym:— Anthoceros dilatatus Steph., Bot. Jahrb. Syst. 8: 95, 1886 ( Stephani 1886). Type:— EQUATORIAL GUINEA. Bioko (Fernando Pó): s. loc., May 1885, Mönkemeyer 15, Stephani herb. no. 20500 (G-00128025! [http://www.ville-ge.ch/musinfo/bd/cjb/chg/adetail. php?id=133424&lang=en]), lectotype here designated [or possibly holotype]). Note:—The selected lectotype was annotated by J. Hasegawa in 1993 as belonging to Anthoceros subg. Folioceros .

Folioceros fuciformis (Mont.) D.C.Bharadwaj, Geophytology 5: 227, 1975 ( Bharadwaj 1975). Basionym:— Anthoceros fuciformis Mont., Ann.Sci.Nat.Bot. (ser.2) 20:296, 1844 [1843] ( Montagne 1844).Type:— RÉUNION.“Bourbon”, 1837, Gaudichaud s.n. (PC-0032647, lectotype selected by Piippo 1993: 36; G-00115646, isolectotype).

= Aspiromitus hildebrandtii Steph., Sp. Hepat. [Stephani] 5: 961, 1916 ( Stephani 1916), syn. nov. Type :— MADAGASCAR. Tananarivo, Hildebrandt s.n. (G-00045056! [http://www.ville-ge.ch/musinfo/bd/cjb/chg/ adetail.php?id=141041&lang=en], lectotype here designated; G-00120707 [http://www.ville-ge.ch/musinfo/bd/ cjb/chg/adetail.php?id=163184&lang=en], PC-0101754, [http://science.mnhn.fr/institution/mnhn/collection/pc/ item/pc0101754], isolectotypes)

Figures

FIGURE. 1. Spores of Anthoceros cristatus. (A) Spores from holotype Stephani herb. no. 24501 (G-00045042) and Aspiromitus dismieri (B). Scale bar is 10 μm. Spores from holotype, Rotereau s.n, Stephanie herb. no.00045054