Orthosyntexis J. Gao, Engel, Shih, & T. Gao, 2021

Gao, Jia, Engel, Michael S., Shih, Chungkun, Ren, Dong & Gao, Taiping, 2021, A new genus of anaxyelid wood wasps from the mid-Cretaceous and the phylogeny of Anaxyelidae (Hymenoptera), Journal of Hymenoptera Research 86, pp. 151-169 : 151

Genus Orthosyntexis J. Gao, Engel, Shih, & T. Gao gen. nov.

Type species.

Orthosyntexis elegans sp. nov.

Etymology.

The new genus-group name is a combination of the Ancient Greek orthós ( ο’ρθός, meaning, “upright” or “erect”, and geometrically "right angle"), and the genus Syntexis (itself from Ancient Greek súntēxis (σύντηξις, meaning, “colliquescence”, “emaciating”, or "wasting away")), type genus of the subfamily Syntexinae . Gender feminine.

Diagnosis.

Female: Antenna with 16 flagellomeres, scape almost 3 × as long as width and twice as long as flagellomere I, flagellomere I nearly 1.5 × as long as flagellomere II. Forewing with pterostigma not enlarged, uniformly sclerotized and of normal width; 1Rs shorter than 1M; cell 1mcu length to width ratio slightly more than 2; 1r-rs and 2rs-m absent; 1Cu obviously shorter than 2Cu; 3Cu shorter than 4Cu; 2m-cu 1.5 × shorter than 1m-cu in forewing; 3rs-m 2 × shorter than 4M. Hind wing with abscissa 2M+Cu; 1M shorter than 2M; m-cu absent, cell r closed. Mesotibia with two apical spurs. Male: Unknown. Immatures: Unknown.

Included species.

Orthosyntexis elegans sp. nov. and O. thanti sp. nov.

Remarks.

Orthosyntexis may be differentiated from Curiosyntexis , in which the forewing pterostigma is desclerotized and 1r-rs is partly reduced ( Kopylov 2019), and from Dolichosyntexis , which has an enlarged pterostigma and forewing 1Cu not obviously shorter than 2Cu ( Kopylov 2019). The new genus can be separated from Parasyntexis by the latter’s narrower pterostigma, narrower than the length of 2r-rs, and 1M longer than 2M in the hind wing ( Kopylov 2019). Orthosyntexis can be distinguished from Sclerosyntexis by the latter’s elongate scape, which is 3 × longer than wide, and the open cell r in the hind wing ( Wang et al. 2020). Orthosyntexis differs from the extant genus Syntexis , which has 3Cu longer than 4Cu and 2m-cu 1.5 × longer than 1m-cu in the forewing ( Rohwer 1915), and from Daosyntexis by 1Rs 3 × being longer than 1M and 3rs-m 2 × longer than 4M in the forewing ( Kopylov et al. 2020). The new genus differs from Cretosyntexis by 2r-rs meeting the pterostigma beyond midlength and Rs+M longer than 2M ( Rasnitsyn and Martínez-Delclòs 2000), and from Eosyntexis in which 1RS is not shorter than 1M ( Rasnitsyn and Martínez-Delclòs 2000).