Anabarhynchus pilbara Ferguson & Irwin

Ferguson, David J., Irwin, Michael E. & Yeates, David K., 2013, New species of Anabarhynchus Macquart (Diptera: Therevidae) from arid and monsoon tropical Australia, Zootaxa 3680 (1), pp. 55-95 : 77-81

Anabarhynchus pilbara Ferguson & Irwin sp. n.

( Figures 17 , 18 , 19 )

Type material. Holotype: Male. AUSTRALIA: Western Australia: Mt. Augustus National Park; malaise at Edney’s Spring; 45m; 25.IV / 7.V.2003. ME Irwin, FD Parker, GPS: 24°21’07”S 116°52’02”E (MEI_173417) ( ANIC). Condition: Ex - alcohol; dried using Hexa Methyl Di Silazane ( HMDS); dorsally pinned with stainless steel pin; scutum cracked; good condition.

Paratypes: 99 Males, 99 Females. AUSTRALIA: Western Australia: 513 52Ƥ, same data as holotype (MEI_173416-173454,173456-173461,173553,173555- 173603,173605) ( ANIC); 403, 41Ƥ, 30 km E Marble Bar at Yandicoogina Creek, malaise nr damp area in sandy wash, M.E. Irwin, F.D. Parker, 212m, GPS: 21°11’00”S 120°01’07”E (MEI_173464, 173465, 173469, 173470-173478, 173480-173548) ( ANIC); 13 30km ESE Three Rivers Station, malaise across dry bed of Gascoyne R., M.E. Irwin, F.D. Parker, 504m, GPS: 25°13’06”S 118°56’09”E (MEI_173464) ( ANIC); 33, 4Ƥ 25km N Marble Bar, malaise across drying pool in Coongan River, M.E. Irwin, F.D. Parker, 136m, GPS: 20°56’02”S 119°51’00”E (MEI_173466-173468, 173549-173552) ( ANIC); 1Ƥ, 295m, 104km E Marble Bar, malaise trap across wash with permanent pools, M.E. Irwin, F.D. Parker, GPS: 21°19’01”S 120°43’03”E (MEI_173604); 23, 11km E Marble Bar at Brockman Creek, malaise in damp, sandy creek bed, M.E. Irwin, F.D. Parker, 187m, GPS: 21°09’00”S 119°51’07”E (MEI_173479, 178022); 23, 260m, 68 km E Marble Bar; malaise trap near pool in small creek bed; M.E. Irwin, F.D. Parker, GPS: 21°13’08”S 120°22’07”E (MEI_173462, 173463); 1Ƥ Millstream-Chichester Nat Park Yarraloola Rd: Fortescue River C. Lambkin T. Weir: ANIC 2119: Malaise in broken creek bank: drying pools GPS: 21°37’38”S 117°07’27”E ( ANIC _29:015371) ( ANIC); 23 Millstream-Chichester Nat Park McKenzie Spring: ANIC 2125 C. L. Lambkin: 299m, malaise over spring in rocky ravine GPS: 21°20’18”S 117°12’38”E ( ANIC _29:015373, 29:015372) ( ANIC).

Addition label on all MEI specimens; (1) CSIRO-Schlinger Pilbara Expedition April 19–May 23, 2003. (2) Anabarhynchus pilbara Irwin. Det. M. E. Irwin, 2010 .

Diagnosis. Frons raised; frontal setae short and pale. Male frons width at anterior ocellus 3x ocellus, female 3.0–4.0x ocellus. Occiput macrosetae pale. Scutum grey pubescence with brown dot-dash mid-dorsal markings. Wing cell m3 open. Postspiracular pile present; katepisternum and prosternal furrow without pile. Forefemur 2 pd, mid-femur 1pd, hind femur, 1 av; all femora pale-yellow.

Description. Male: Body length: 8.5mm. Wing length: 7mm. Head. Integument black. Frons raised; frons width at anterior ocellus 3x ocellus; ocellar tubercle flat; antennal base positioned low on frons and slightly protruding; parafacials brownish grey, frons brown grey; middle frons with dark grey upwardly curved mark meeting at middle, varies in size depending on angle of view; upper and middle frons with several lines of short depressed dark setae, lower frons with single pale short appressed setae less than quarter length of scape. Scape length 2x width; scape and pedicel brownish grey, 1st segment of flagellum basally pale brown, rest dark brown with a few short black setae to dorsal basal third, flagellar style blackish-brown, half length of flagellum. Occiput convex with yellowish-grey pubescence, several indistinct rows of pale macrosetae 35 each side; postocciput area to gena grey with dense, long, pale, hair-like pile. Palpus whitish, with sparse pale hair-like pile; labellum pale brown, prementum without dark setae. Thorax. Integument generally black; postpronotal lobe, upper katepisternum, anterior meron anepimeron, katatergite and coxae integument yellow brown. Scutal chaetotaxy black (pairs): np, 4; sa, 2; pa, 1; dc, 2; sc, 2. Scutum with thin brown dorsal line within a broader grey band, margined by paler grey lines, brown marks punctuate the perimeter the raised areas pre and post-transverse suture; scutal surface with short, sparse, appressed, pale setae. Postspiracular pile present; katepisternum and prosternal furrow without pile; pleuron with thick grey pubescence; coxae yellow-grey pubescence with long pale setae admixed strong black macrosetae. Wing. Cell m3 open; weakly hyaline; pale brown tint; veins brown; stigma pale brown; costal setae beyond humeral cross-vein biserially arranged. Haltere. Pedicel buff-white; scabellum white. Legs. Forefemur with 2 pd towards middle; mid-femur 1 pd; hind femur 1–2 av; all femora pale-yellow with thin pale yellow-grey pubescence to ventral surfaces; all femora with sparse distribution of short semi-erect pale pile; admix with short black setae predominantly to dorsal apical half; all femora with sparse black setae admix with pale appressed pile dorsally; ventral pile erect, less than half the width of hind femoral; tibias and basitarsus yellow-brown darkening apically. Abdomen. Integument mottled yellow-brown; tergite anterior bands yellowbrown without pubescence with sparse appressed black pile; posterolaterally grey with appressed white setae; tergites 2–6 posterior marginal bands to when viewed posteriorly appear bright white, viewed anteriorly appear grey; sternites yellow-brown thin grey pubescence with sparse pale semi-erect pile; epandrium yellow-grey with yellowish setae; gonocoxites yellow-brown with thin grey pubescence; line of bright white pubescence along ventromedial plane. Terminalia. Paratype male (MEI_173448): epandrium ( Fig. 19 a), one and three quarters times wide as long, posterior lateral corner rounded ventrally with sclerotised ridge to ventral surface; the centreline of sub-epandrium plate ridged. Gonocoxite ( Fig. 19 b), semi-spherical slightly longer than wide when viewed ventrally, middle posterior ventral edge with small lobe shaped outer gonocoxal process. Joined along hypandrium. Gonocoxal apodeme elongated darkly sclerotised. Inner gonocoxal process longer than gonostylus, ventrally directed sub-apical setae on inner edge; gonostylus elongate directed dorsally with several setae on inner dorsal surface, apical end reflex dorsally, several elongate setae along sub-apical lateral edge. Ventral lobe absent. Aedeagus ( Fig. 19 c, d): distiphallus strongly curved ventrally. Parameral sheath sclerotised. Dorsal apodeme of parameral sheath broadly triangular, medially thinly sclerotised; apical ends darkly sclerotised ventrally and inwardly directed. Ventral apodeme basally narrow boarding apically triangular. Lateral ejaculatory apodeme elongated triangular, band-like. Ejaculatory apodeme cylindrical with sub-apical lobes.

Variation. Male. Body length range: 7–8.5mm. Wing length range: 5.5–7.5mm. Frons width at anterior 2.5– 3.1x ocellus; occiput macrosetae 25–35 each side. Forefemur, 1–4 pd; mid-femur, 1 pd, 0–2 pv and 0–2 av; hind femur, 1–2 av. Female. Body length: 7.5–9.5mm. Wing length: 6.5–8mm. Frons width at anterior ocellus 3.0–4.0x ocellus; frons with brown markings slightly more extensive than in male. Anterior bands to tergite 2–6 darker brown. Paratype female (MEI_173605): sternite 8 ( Fig.19 f), oval in shape; darkly sclerotised; anterior margin deep concaved; posterior with board rounded lobe inserted into main sclerite; elongate slightly depressed area along mid-line; long black setae either side of centre line. Furca ( Fig. 19 e), oval shaped; posterior frame thin; middle with long low lobe latererally directed; lateral edge of anterior beam with anterior projections; anteroventral lobe absent.

Etymology. The specific epithet ‘ pilbara ’ is derived from the geographic location where the type specimens were collected.

Comments. Described from a large series of specimens collected in April and May during the CSIRO- Schlinger Pilbara Expedition that have a wide distributed across the Pilbara area of north west Western Australia.

Keys to couplet 113 in Lyneborg (2001) and appearing similar to A. tener Lyneborg both with pale occiput macrosetae; A. pilbara sp. n., are all pale, while A. tener are admix pale and black. Frontal pile in A. pilbara is very short and pale concentrated to the mid and upper frons, with only several above the antenna sockets; while A. tener are pale admix with black, short and erect sparsely reach lower frons. Femoral setae on A. pilbara are forefemur, 2 pd, 0–1av; mid-femur 1 pd; hind femur 1–2 av. While femoral setae on A. tener are forefemur, 1–2 pd, 1 pv; midfemur 1 pd, 2 pv; hind femur, 1–2 av, 1–2 pv. Abdominal colouration in A. pilbara is yellowish, while A. tener is of a grey appearance.

Anabarhynchus pilbara sp. n., closely related to A. gascoyne Lyneborg of the kroeberi species-group, sharing a similar simple furca. Sternite 8 is elongate with depressed midline and is placed with the kroeberi species-group. Related to both A. ewamin sp. n., A. ravenshoensis sp. n., and A. tribulationensis sp. n., for femoral macrosetae differences of the kroeberi species-group see Table 1.

Figures

FIGURE 17. Anabarhynchus pilbara sp. n., holotype male, (MEI _ 173417): lateral view.

FIGURE 18. Anabarhynchus pilbara sp. n., holotype male, (MEI _ 173417): oblique view.

FIGURE 19. Anabarhynchus pilbara sp. n., paratype male terminalia (MEI _ 173448): A, epandrium, left dorsal view, right ventral view; B, gonocoxite left dorsal view, right ventral view; C, aedeagus, dorsal view; D, aedeagus, lateral view; paratype female (MEI _ 173605); E, furca; F, female sternite 8, ventral surface. Scale line = 0.5 mm.