Padillothorax mulu Maddison, sp. nov.

Maddison, Wayne P., Beattie, Imara, Marathe, Kiran, Ng, Paul Y. C., Kanesharatnam, Nilani, Benjamin, Suresh P. & Kunte, Krushnamegh, 2020, A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini), ZooKeys 1004, pp. 27-97 : 27

Padillothorax mulu Maddison, sp. nov. Figs 18 , 25 , 176-187

Type material.

All from Malaysia: Sarawak: Mulu Nat. Pk., in UBCZ. Holotype male (specimen IDWM.20009) from Summit Trail near Camp 1, 4.0496°N, 114.8589°E to 4.0493°N, 114.8594°E, 220 m el. 21 March 2012 Maddison/Piascik/Ang WPM#12-067. Paratype female (specimen SWK12-EP0105) from Summit Trail near Camp 1, 4.0489°N, 114.8606°E to 4.0486°N, 114.8610°E, 280 m el. 21 March 2012 Maddison/Piascik/Ang WPM#12-071. Paratype male (specimen SWK12-2556) from Summit Trail near Camp 1, 4.0491°N, 114.8601°E to 4.0489°N, 114.8606°E, 270 m el. 21 March 2012 Maddison/Piascik/Ang WPM#12-070. Paratype female (specimen SWK12-EP0108) and male (specimen IDWM.20010) from Base Trail near Camp 1, 4.0543°N, 114.8534°E to 4.0544°N, 114.8531°E, 130 m el. 23 March 2012 Maddison/Piascik/Ang WPM#12-084.

Etymology.

From the name of the type locality. Other names: In Maddison (2015b) and WPM’s field or lab notebooks this species was grouped with P. badut under the informal name “BVBND”, until it was distinguished from that species as “BVBND-M”.

Diagnosis.

Very similar to P. badut , differing most notably in details of genitalia. The embolus of P. mulu is shorter, and has a toothed ridge on the prolateral base; the epigyne has the openings exposed on a more or less flat surface.

Description.

Male (holotype, specimen IDWM.20009). Carapace length 2.5; abdomen length 3.5. Colour and structure matches that of P. badut as described above. Cheliceral teeth not examined. Palp pale except for base of femur. Base of embolus with various fine teeth; embolus shorter than in P. badut (Fig. 176 ).

Female (paratype, specimen SWK12-EP0105). Carapace length 2.1; abdomen length 3.3. Colour and structure matches that of P. badut as described above. Cheliceral teeth not examined in this specimen, but another female from Mulu has five retromarginal teeth. Epigyne (Fig. 178 ) similar to that of P. badut , except for the openings being exposed, not under a fold.

Natural history.

As for P. badut , on large-leaved understory plants. They often hold the front legs out or to the front.

Additional material examined.

All from Malaysia: Sarawak: Mulu Nat. Pk., in UBCZ. One female (specimen SWK12-EP0107) from Base Trail near Camp 1, 4.0543°N, 114.8534°E to 4.0544°N, 114.8531°E, 130 m el. 23 March 2012 Maddison/Piascik/Ang WPM#12-084. One female (specimen SWK12-2574) from Summit Trail near Camp 1, 4.0491°N, 114.8601°E to 4.0489°N, 114.8606°E, 270 m el. 21 March 2012 Maddison/Piascik/Ang WPM#12-070.

Figures

Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20).

Figures 176 - 187. Padillothorax mulu sp. nov. 176 male left palp, ventral view (holotype IDWM. 20009) 177 same, retrolateral view of tibia 178 epigyne, ventral (specimen SWK 12 - EP 0105) 179 vulva, dorsal 180 - 183 Male (SWK- 2556) 184 - 187 female (SWK 12 - 2574). Scale bars: on genitalia 0.1 mm; on bodies 1.0 mm.