Stantonia sommeijeri, Braet & Quicke, 2004

Braet, Yves & Quicke, Donald, 2004, A phylogenetic analysis of the Mimagathidini with revisionary notes on the genus Stantonia Ashmead, 1904 (Hymenoptera: Braconidae: Orgilinae), Journal of Natural History 38 (12), pp. 1489-1589 : 1580-1583

Stantonia sommeijeri sp. n.

( figures 48 , 57–59 )

Etymology. From the name of the collector.

Type material. HOLOTYPE, ” ( RMNH): Tobago: Goldborough , 23 February– 24 March 1994 (M. J. Sommeijer) . PARATYPES: Brazil ( AEI): Para : Jacareacanga: ” and „, December 1968 (M. Alvarenga) . Colombia ( IAVH): Amazonas : one specimen unsexed, PNN, Hamacayacu Caño Mata-Mata, February 1989 (Malaise trap; M. Kelsey)— Stantonia sp.—IAVHHB508 . Costa Rica ( TAMU): ” and „, 11 mi. from Turrialba , ‘ Los Esperales’, C.A. T.I.E., 5 February 1985 (P. Stansly) . Peru

(AEI): Avispas: ” and „, nr Maracapata, 30 m, 1–15 October 1962 (L. Peña); „, idem but: ‘ 1–15 October 1962, collection R. D. Shenefelt’. Loromayo: „, 4–10 September 1962 (L. Peña). Trinidad ( USNM): „, Rio, Pan, 29 March 1912 (A. Busck). Venezuela ( MIZA): Cojedes, El Pao: „, Higuerotal, 24–31 June 1995 (9 ‡ 49’27@N, 68 ‡ 2’50@W, 376 m, Malaise trap, Proyecto MIZA-DHC, A. Alemán); „, Galeras, 19–21 December 1994 (9 ‡ 34’24@N, 68 ‡ 09’7@W, 220 m, Proyecto MIZA- DHC, A. Alemán) .

Holotype, ”, length of body 4.56 mm, of fore wing 4.8 mm.

Head ( figure 59 ). Antenna with 49 segments, twice as long as fore wing, length of third segment 1.25 times fourth, length of third, fourth and penultimate segment 5, 4 and 1.6 times their maximal width, apical segment three times its maximal width; maxillary palp 1.1 times height of head; length of eye in dorsal view three times temple; eyes with sparse and short setae; temple directly narrowed posteriorly, coriaceous ventrally and smooth dorsally; OOL: diameter of ocellus: POL~ 6.0:4.0:3.0; frons smooth medially; vertex convex, slightly punctate, flattened near stemmaticum; face and clypeus convex and punctate; length of malar space one times basal width of mandible; occipital flange weakly rounded posteriorly and straight ventrally; malar suture present

Mesosoma ( figure 57 ). Length of mesosoma 1.3 times its maximal height; side of pronotum punctate posteriorly, smooth near tegulae, medially crenulate and remainder smooth; mesopleuron finely punctate beside precoxal sulcus; precoxal sulcus weakly crenulate anteriorly and reaching anterior border of mesopleuron; metapleuron shiny and punctate; mesoscutum sparsely punctate, depressed posteriorly at junction of notauli, lateral lobes more deeply punctate; scutellum smooth; scutellar sulcus with small carinae posteriorly; surface of propodeum medially with weak transverse rugae, smooth laterally ( figure 58 ); metapleural flange small and angulate/acute.

Wings. Fore wing: r: SR1z3-SR~9:40; 1-SRzM straight basally joined to 1-M; second submarginal cell strongly petiolate; subbasal cell mainly glabrous anteriorly (only three to four setae near cu-a vein); vein 3-CU1 as long as vein CU1b. Hind wing: 2-M coloured; membrane largely glabrous basally.

Legs. Hind coxa with several coarse rugae dorsally ( figures 48 , 58 ), punctategranulose dorsally; lengths of hind femur, tibia and basitarsus 4.8, 8.3 and 7.3 times their maximal widths, respectively; hind femur punctate; tibial spurs 0.6 and 0.36 times basitarsus; hind tibia with three apical teeth.

Metasoma. First tergite 3.3 times its apical width; flattened surface of first tergite smooth to finely coriaceous and shiny ( figure 48 ); spiracles weakly protruding; second tergite smooth and shiny; T2 1.2 times longer than T3; second suture smooth, straight; ovipositor sheath less than 0.1 times fore wing length.

Colour. Yellow-brownish. Tergites except third, propodeum dorsally, mesoscutum (except mid-longitudinal part of median lobe), vertex, upper part of temple, outer stripe of scapus, hind coxa dorsally, hind tibia apically, hind tibial spurs, apical patch (occupying half of marginal cell) of fore wing, dorsal part of outer side of hind femur blackish; tegula, humeral plate, mid tibia apically, inner side of hind femur, hind trochantellus, hind tibia basally, mid and hind tarsus except basitarsus, frons, antennal segments brownish; extreme base of T1, apical transverse band of propodeum, hind basitarsus whitish yellow.

Variation. Frons sometimes black, scutellum black to yellowish, metanotum black only medially to completely, mesopleuron antero-ventrally dark brownish to yellowish, mid femur yellowish to orange-brownish on inner side, only T1 and T2 sometimes dark brownish with remaining tergites marked with pale brown spots, mid tibia completely brownish, hind tibia and hind basitarsus completely black. Number of antennal segments varies between 46 and 51.

Stantonia spasskensis Belokobylskij, 1993 Stantonia spasskensis Belokobylskij, 1993: 97 .

Material examined ( ZIRAS). Vietnam: ”, pr. Ha Son Binh , Da Bac, Tuly, 22 October 1990 (bamboo forest; S. Belokobylskij) .

Remarks. This female is largely as the male holotype except for the following characters: antenna with 48 segments, median segments (around the 17th) whitish, wings weakly infuscate especially apically, scutellar sulcus with one mid-longitudinal carina and several smallest posteriorly, surface of propodeum smoothpunctate, surface of hind coxae aciculate to coriaceous-granulose, sculpture of propodeum and hind coxae weak but parallel, mid-longitudinal carina of propodeum absent, length of ovipositor sheath 0.8 times fore wing, hind tarsus whitish except one faint basal dark stripe on basitarsus. Occurs in the East Palaearctic in Russia (Primorskiy Kray).

Stantonia sumatrana Enderlein, 1908

( figures 102, 104 )

Stantonia sumatrana Enderlein, 1908: 110 ; Shenefelt, 1970: 268; van Achterberg, 1987: 47 [examined].

Type material. HOLOTYPE, ” ( PAIZ): Sumatra: Soekaranda, Dr H. Dohrn S. — Type— Stantonia sumatrana n. sp. ”—Mus. Zool. Polonicum, Warszawa, 12/45.

Other material examined. Australia ( ANIC): N. Queensland: ”, Kuranda, 21 June–24 July 1913 (1100 feet; R. E. Turner; 1913-438); ”, Mossman Gorge, 23 April 1967 (D. H. Cohen); ” and „, Babinda, The Boulders, 10 May 1967 (D. H. Cohen); „, Cape York pen., Iron Range, 1–9 June 1971 (S. R. Monteith). N. T.: „, Katherine, 7 June 1964 (K. R. Morris); ”, 1 km N of Cahills X-ing, East Alligator River , 7–8 June 1973 (12 ‡ 25’S, 132 ‡ 58’E; J. C. Cardale) . Malaysia and Singapore ( USNM): Singapore: 8”” and 6„„. Selangor: Kuala Lumpur: 2””, 1950 (Army Scrub. Typhus Unit). Penang Is. [~ Pulau Pinang]: ” and 3„„. Philippines ( USNM): Mindanao: Surigao, 2”” and 4„„; Iligan: ” .

Holotype, ”, length of body 8 mm, of fore wing 7.4 mm.

Head ( figure 102 ). Remaining antennal segments 27 and 18, length of third segment 1.2 times fourth, length of third and fourth segments 3.7 and 3 times their maximal width; eye glabrous; length of eye in dorsal view 2.4 times temple; temple coriaceous, directly narrowed posteriorly, striate dorsally (near eye); OOL: diameter of ocellus: POL~7.0:4.0:2.0; frons smooth medially; vertex convex, flattened near stemmaticum; face flattened in lateral view, smooth; clypeus flattened in lateral view, smooth and shiny; but both clypeus and face convex; length of malar space equal to basal width of mandible; malar suture present; occipital flange rounded.

Mesosoma ( figure 104 ). Length of mesosoma 1.3 times its maximal height; side of pronotum medially crenulate, punctate ventrally, and remainder smooth; mesopleuron smooth and shiny between punctures; precoxal sulcus crenulate medially, punctate elsewhere; metapleuron punctate; mesoscutum sparsely punctate; scutellum convex; propodeum medially convex in lateral view, smooth with transverse irregular rugae posteriorly, laterally with sparse punctures; metapleural flange angled.

Wings. Fore wing: r: SR1z3-SR~13:53; 2-SR: r-m~18:11; second submarginal cell subsessile; subbasal cell glabrous anteriorly (near MzCU1 vein); vein 3-CU1 twice longer than vein CU1b. Hind wing: 2-M coloured; membrane largely glabrous basally.

Legs. Hind coxa punctate with few rugae dorsally; hind femur coarsely punctate; lengths of hind femur, tibia and basitarsus 5.2, 6.4 and 8.7 times their widths, respectively; tibial spurs 0.46 and 0.38 times basitarsus; hind tibia with three apical teeth.

Metasoma. First tergite weakly trapezoidal, 2.7 times longer than apically wide, smooth and shiny, apically weakly coriaceous, medially concave and apically weakly convex in lateral view; spiracles protruding; second tergite weakly punctatecoriaceous; T 2 1.38 times longer than T 3; second suture smooth, straight; ovipositor sheath less than 0.1 times fore wing length.

Colour. Brownish yellow. Telotarsi, apical patch of hind tibia, ovipositor sheath blackish; dorso-apical patch of hind femur, apical patch of mid tibia, hind basitarsus (one-fifth apical) and rest of hind tarsi, metasoma brownish; wings, antenna apically and dorsally, stripe on outer side of scapus infuscate; humeralplate brownish infuscate.

Figures

FIGS 47–52. (47, 50) Stantonia mazatlanensis n. sp., holotype, ”. (48) S. sommeijeri n. sp., holotype, ”. (49) S. hammersteini, holotype, ”. (51) S. maracapatana n. sp., holotype, „. (52) S. sirena n. sp., paratype, ”. Detail of first tergite, lateral view (47, 48) and dorsal view (49, 50); detail of outer claws of hind leg (51); detail of T2–T3 suture in dorsal view (52).

FIGS 53–58. (53, 55, 56) Stantonia sirena n. sp., paratype, ”. (54) S. rufithorax, holotype, „. (57, 58) S. sommeijerin. sp., holotype, ”. Head inlateral view (53, 54); first tergite in lateral (55) and dorsal view (56); mesosoma in lateral view (57); propodeum and T1 (58).

FIGS 59–64. (59) Stantonia sommeijeri n. sp., holotype, ”. (60–63) S. pachycornis n. sp., holotype, ”. (64) S. mazatlanensis n. sp., holotype, ”. Head in lateral view (59, 60); mesosoma in lateral view (61, 64); propodeum and first tergite in lateral (62) and dorsal view (64).

FIGS 101–106. (101, 103) Stantonia maracapatana n. sp., holotype, „. (102, 104) S. sumatrana, holotype, ”. (105) S. veronicae n. sp., paratype, „. (106) S. flava, holotype, ”. Head in lateral view (101, 102); mesosoma in lateral view (103, 104, 106); propodeum (105).