Epimetopus angulatus Balfour-Browne, 1949

Perkins, Philip D., 2012, 3531, Zootaxa 3531, pp. 1-95 : 43-45

Epimetopus angulatus Balfour-Browne, 1949

Figs. 78 (habitus), 79, 80 (aedeagus), 126 (map)

Epimetopus angulatus Balfour-Browne 1949: 16 .

Epimetopus angulatus: Oliva (1986: 8) ; misidentification.

Epimetopus angulatus: Fikáček , et al. (2011: 489).

Type material: Holotype (male): " Type [red disc]; Yungas Valley. Bolivia. 12–13.vi.1937; H. E. Hinton Collector; Epimetopus angulatus Type! J. Balfour-Browne det.; Hinton Coll. B.M. 1939-583" ( BMNH).

Differential Diagnosis. The dorsal habitus of E. angulatus is very similar to that of E. peruvianus ; E. angulatus is slightly larger (ca. 2.73 vs. 2.62 mm) but otherwise very similar ( Figs. 78, 82). Reliable determinations will be based on examination of the aedeagi of the two species, which are markedly dissimilar ( Figs. 78, 80, 82). The aedeagus could be compared with that of E. coleuncus ( Fig. 85), but differs from that in several respects, including the following: 1) the aedeagus is wider, the lateral processes of the median lobe (the part contacting the parameres) being notably larger; 2) the median lobe apical part is narrower and is emarginate laterally, whereas in E. coleuncus the apical part is wider and arcuate laterally, 3) the hooks rest in a slot on each side of the median lobe, whereas in E. coleuncus the hooks each rest in a groove on the dorsal surface of the median lobe; 4) the parameres are shaped differently; and 5) the basal piece is longer in E. coleuncus . The aedeagus of E. angulatus differs markedly from E. mendeli in overall shape, and many details.

Description. Size: holotype (length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.73/1.43; head (width) 0.68; pronotum 0.95/1.00; elytra 1.82/1.43. Habitus as illustrated ( Fig. 78). Head and pronotum dark brown to piceous; elytra brown to dark brown, without fascia; maxillary palpi light brown; venter and coxae piceous, remainder of legs reddish brown. Eye canthus incomplete, ca. 3 facets between canthus and posterior margin. Pronotum with hood carinae slightly arcuate, confluent at anterior tip, grooved throughout; primary lateral lobe with anterior angle produced and acute; secondary lateral lobe extremely small or absent. Elytron with four granulate carinae, third interrupted anteriorly for distance of ca. 3–4 punctures; punctures round, small, each with moderately large granule at anterior margin, non-carinate intervals with irregular row of granules. Protibiae straight or very slightly emarginate on inner margin, outer margin weakly arcuate. Metaventral depression small, ca. 8 granules along base.

Distribution. Currently known only from the type locality in Bolivia ( Fig. 126).

Notes. Oliva (1986; fig. 4), illustrated an aedeagus supposedly of E. angulatus . However, it is not, and appears instead to perhaps be the aedeagus of E. coleuncus . The aedeagus of the holotype of E. angulatus was illustrated by Fikáček et al. (2011), and a detail of the apex of the same aedeagus is illustrated herein ( Fig. 79). Because of the error, and the fact that other species have a habitus similar to that E. angulatus , the distribution records given by Oliva (1986) must be considered invalid until being confirmed by the dissection of males.

Epimetopus mendeli Fiká č ek, Barclay & Perkins, 2011

Fig. 126 (map)

Epimetopus mendeli Fikáček, Barclay & Perkins, 2011: 480 .

New collection records: Peru: Cusco, Pil [l]copata, premontane moist forest, elev. 600 m, 13° 4' S, 71° 10' W, 8–10 xii 1979, J. B. Heppner (1 USNM) ; same locality, elev. 600 m, 13° 4' S, 71° 10' W, 11–14 xii 1979, J. B. Heppner (2 USNM) ; Prov. Quispicanchis, Quincemil [Quince Mil], 13° 14' S, 70° 46' W, 6–11 x 1976, Robert Gordon (1 USNM) ; San Pedro (at km 152), MV light, elev. 1430 m, 14° 11' S, 71° 20' W, 30 viii 1989, R. A. Faitoute et al. (8 USNM) ; Santa Isabel, Cosnipata Valley , 13° 0' S, 71° 18' W, 9 xii 1951, Felix Woytkowski (1 FMNH) ; Madre de Dios, Hostel Erica (near Salvacion ), elev. 550 m, 12° 53' S, 71° 14' W, 3–5 ix 1989, R. A. Faitoute et al. (1 USNM) .

Differential diagnosis. The dorsal habitus is similar to that of E. angulatus , but E. mendeli is considerably larger (ca. 3.10 vs. 2.73 mm), and has more elongate elytra (l/w ratio ca. 1.40 vs. 1.26), and a distinctive elytral fascia (see Figs. in Fikáček et al., 2011). The aedeagus of E. mendeli differs markedly from E. angulatus in overall shape, and many details ( Figs. 79, 80). The aedeagus of E. mendeli is somewhat similar to that of E. coleuncus . It differs from E. coleuncus in the following: 1) the shape and length of the parameres, which are shorter and lack the subapical lateral sinuations; 2) the shape of the apical part of the median lobe (more sharply rounded at the tips); and 3) in having narrower lateral processes of the median lobe.

Distribution. Currently known only from Peru ( Fig. 126).

Notes. For a detailed description see Fikáček et al. 2011.

Epimetopus flavicaptus Fiká č ek, Barclay & Perkins, 2011

Figs. 81 (habitus), 126 (map)

Epimetopus flavicaptus Fikáček, Barclay & Perkins, 2011: 488 .

Type material: Holotype (male): Ecuador, Napo Province [see data in Fikáček, et al. 2011].

Differential diagnosis. Distinguished from other members of the Mendeli group by the transverse or slightly emarginate anterior margin of the pronotal hood ( Fig. 81). The aedeagus is illustrated by Fikáček et al. (2011). The distal part of the median lobe is clearly diagnostically different than that of the other members of the Mendeli group.

Distribution. Currently known only from the type locality in Ecuador ( Fig. 126).