Pelecyphora vivipara (Nutt.) D.Aquino & Dan. Sanchez, 2022

Sanchez, Daniel, Vazquez-Benitez, Balbina, Vazquez-Sanchez, Monserrat, Aquino, David & Arias, Salvador, 2022, Phylogenetic relationships in Coryphantha and implications on Pelecyphora and Escobaria (Cacteae, Cactoideae, Cactaceae), PhytoKeys 188, pp. 115-165 : 115

Pelecyphora vivipara (Nutt.) D.Aquino & Dan. Sanchez comb. nov.

Cactus viviparus ≡ Cactus viviparus Nutt. Nutt., Cat. Pl. Upper Louisiana no. 22. 1813. Mammillaria vivipara (Nutt.) Haw., Suppl. Pl. Succ.: 72. 1819. Echinocactus viviparus Poselg., Allg. Gartenzeitung 21: 107. 1853. Mammillaria radiosa var. vivipara (Nutt.) Schelle, Handb. Kakteenkult.: 236. 1907. Coryphantha vivipara (Nutt.) Britton & Rose, Ill. Fl. N. U.S.: 571. 1913. Escobaria vivipara (Nutt.) Buxb., Oesterr. Bot. Z. 98: 78. 1951. Coryphantha neovivipara Y. Itô, Cactaceae : 556. 1981. comb. inval. Type: United States, North Dakota, McClean County, 12 mi [19.31 km] E of Fort Mandan, E of Missouri River, Jun 1971, L. Mitich s.n. (neotype, designated by Mitich & Benson, Cact. Succ. J. (Los Angeles): 8. 1977: POM [317948]; isoneotype NDA).

Mammillaria radiosa = Mammillaria radiosa Engelm., Boston J. Nat. Hist. 6: 196. 1850. Echinocactus radiosus Poselg., Allg. Gartenzeitung 21: 107. 1853. Mammillaria vivipara var. radiosa Engelm. Proc. Amer. Acad. Arts 3: 269. 1856. Mammillaria vivipara subsp. radiosa Engelm., Rep. U.S. Mex. Bound. Cact.: 15. 1858. Cactus radiosus (Engelm.) J.M.Coult., Contr. U.S. Natl. Herb. 3: 120. 1894. Coryphantha radiosa (Engelm.) Rydb., Fl. Rocky Mts.: 581. 1917. Neomammillaria radiosa (Engelm.) Rydb., Fl. Plains N. Amer.: 562. 1932. Coryphantha vivipara var. radiosa (Engelm.) Backeb., Cactaceae 5: 2998. 1961. Escobaria vivipara var. radiosa (Engelm.) D.R.Hunt, Cact. Succ. J. Gr. Brit. 40: 13. 1978. Escobaria radiosa (Engelm.) G.Frank place of publication unknown, nom. inval. Coryphantha neovivipara var. radiosa (Engelm.) Y. Itô, Cactaceae : 556. 1981. Type: United States, Texas, sterile soils on the Pierdenales [Pedernales], and cult. In St Louis, Jun 1846, F. Lindheimer s.n. (lectotype, designated by Benson, Cacti U. S. Canada: 960. 1982: MO [2017377 image!, 2017376 image!]).

Mammillaria vivipara vera = Mammillaria vivipara var. vera Engelm., Proc. Amer. Acad. Arts 3: 269. 1856. Type: United States. On the upper Missouri and Yellowstone rivers (Not preserved).

Mammillaria vivipara radiosa neomexicana = Mammillaria vivipara var. radiosa subvar. neomexicana Engelm., Proc. Amer. Acad. Arts 3: 269. 1856. Cactus radiosus var. neomexicanus (Engelm.) J.M.Coult., Contr. U.S. Natl. Herb. 3: 120. 1894. Cactus neomexicanus (Engelm.) Small, Fl. S.E. U.S.: 812. 1903. Mammillaria neomexicana (Engelm.) A.Nelson, New Man. Bot. Centr. Rocky Mt.: 327. 1909. Coryphantha neomexicana (Engelm.) Britton & Rose, Cactaceae 4: 45. 1923. Escobaria neomexicana (Engelm.) Buxb., Oesterr. Bot. Z. 98: 78. 1951. Coryphantha vivipara var. neomexicana (Engelm.) Backeb., Cactaceae 5: 2999. 1961. Coryphantha neovivipara var. neomexicana (Engelm.) Y. Itô, Cactaceae : 556. 1981. nom. inval. Escobaria vivipara var. neomexicana (Engelm.) Buxb., Kakteen (H. Krainz) 108c, 1973. Type: United States, South New Mexico, 1849, C. Wright s. n. (lectotype, designated by Benson, Cacti U. S. Canada: 960. 1982: MO [2019650 image!]).

Mammillaria arizonica = Mammillaria arizonica Engelm., Bot. California 1: 244. 1876. Cactus radiosus var. arizonicus (Engelm.) J.M.Coult., Contr. U.S. Natl. Herb. 3: 121. 1894. Mammillaria radiosa var. arizonica (Engelm.) K.Schum., Gesamtbeschr. Kakt.: 481. 1898. Mammillaria radiosa f. arizonica (Engelm.) Schelle, Handb. Kakteenkult.: 235. 1907. Coryphantha arizonica (Engelm.) Britton & Rose, Cactaceae 4: 45. 1923. Mammillaria vivipara var. arizonica (Engelm.) L.D.Benson, Proc. Calif. Acad. Sci., ser. 4, 25: 263. 1944. Coryphantha vivipara var. arizonica (Engelm.) W.T.Marshall, Desert. Bot. Gard. Arizona, Sci. Bull. 1: 94. 1950. Escobaria arizonica (Engelm.) Buxb. in Oesterr. Bot. Z. 98: 78. 1951. Escobaria vivipara var. arizonica (Engelm.) D.R.Hunt, Cact. Succ. J. Gr. Brit. 40: 13. 1978. Coryphantha neovivipara var. arizonica (Engelm.) Y. Itô, Cactaceae : 556. 1981. Nom. inval. Type: United States, Arizona, E. Coues & E. Palmer s. n. (lectotype, designated by Benson, Cacti U. S. Canada: 961. 1982: MO [2017352 image]).

Mammillaria deserti = Mammillaria deserti Engelm., Bot. California 2: 449. 1880. Cactus radiosus var. deserti (Engelm.) J.M.Coult., Contr. U.S. Natl. Herb. 3: 121. 1894. Mammillaria radiosa var. deserti (Engelm.) K.Schum., Gesamtbeschr. Kakt.: 481. 1898. Mammillaria radiosa f. deserti (Engelm.) Schelle, Handb. Kakteenkult.: 236. 1907. Coryphantha deserti (Engelm.) Britton & Rose, Cactaceae 4: 46. 1923. Mammillaria vivipara var. deserti (Engelm.) L.D.Benson in Proc. Calif. Acad. Sci., ser. 4, 25: 263. 1944. Coryphantha vivipara var. deserti (Engelm.) W.T.Marshall, Desert. Bot. Gard. Arizona, Sci. Bull. 1: 94. 1950. Escobaria deserti (Engelm.) Buxb., Oesterr. Bot. Z. 98: 78. 1951. Coryphantha chlorantha var. deserti (Engelm.) Backeb., Cactaceae 5: 3003. 1961. Escobaria vivipara var. deserti (Engelm.) D.R.Hunt, Cact. Succ. J. Gr. Brit. 40: 13. 1978. Type: United States, California, at Ivapah, 30 miles northeast of San Bernardino, in one mountain range stretching into the desert, S. B. Parish 455 (lectotype, designated by Benson, Cacti U. S. Canada: 961. 1982: MO [2267169 image!]).

Mammillaria hirschtiana = Mammillaria hirschtiana F.Haage, Monatsschr. Kakteenk. 6: 127. 1896. Type: No designated.

Mammillaria radiosa texensis = Mammillaria radiosa var. texensis Schelle, Handb. Kakteenkult.: 236. 1907. Type: No designated.

Mammillaria ramosissima = Mammillaria ramosissima Quehl, Monatsschr. Kakteenk. 18: 127. 1908. Type: United States, California, R. C. Orcutt s. n. (lectotype, designated here, Monatsschr. Kakteenk.: 127. 1908: Illustration " Mamillaria ramosissima Quehl Nach einer von Herr De Laet aufgenommenen Photographie").

Coryphantha bisbeeana = Coryphantha bisbeeana Orcutt, Cactography: 3. 1926. Escobaria bisbeeana (Orcutt) Borg, Cacti: 305. 1937. Coryphantha vivipara var. bisbeeana (Orcutt) L.D.Benson, Cacti Ariz. ed. 3: 25. 1969. Escobaria vivipara var. bisbeeana (Orcutt) D.R.Hunt, Cact. Succ. J. Gr. Brit. 40: 13. 1978. Type: United States, Arizona, J. N. Rose 11958 (lectotype, designated by Benson, Cacti Ariz. ed 3: 25. 1969: US [3050430 image!]).

Coryphantha columnaris = Coryphantha columnaris Lahman, Cact. Succ. J. (Los Angeles) 6: 27. 1934. Type: United States, Oklahoma, Jackson County, near Altus, 600 ft [182 m], 1926, M. S. Lahman s.n. (holotype: MO).

Coryphantha fragrans = Coryphantha fragrans Hester, Desert Pl. Life 13: 152. 1941. Type: United States, Texas, in a fertile, sandy loam valley, along the railroad right-of-way and Highway 90, a few miles west of Sanderson, 03 May 1940, J. P. Hester s.n. (holotype: DS [278622 image!]).

Coryphantha rosea = Coryphantha rosea Clokey, Madroño 7: 75. 1943. Coryphantha vivipara var. rosea (Clokey) L.D.Benson, Cacti Ariz. ed. 3: 26. 1969. Escobaria vivipara var. rosea (Clokey) D.R.Hunt, Cact. Succ. J. Gr. Brit. 40: 13. 1978. Type: United States, Nevada, Clark County, between Kyle Canyon and Deer Creek. 24 Jun 1938, I. W. Clokey 8038 (holotype: UC [905407 image!]; isotypes: F [52864 image!]; MEXU [86081 image!]; MICH [1127565 image!]; NY [120673 image!, 120672 image!], TEX [255617]).

Coryphantha oklahomensis = Coryphantha oklahomensis Lahman, Cact. Succ. J. (Los Angeles) 21: 165. 1949. Escobaria oklahomensis (Lahman) Buxb., Oesterr. Bot. Z. 98: 78. 1951. Type: United States, Oklahoma, Caddo County, Range throughout western Oklahoma, collector not mentioned (lectotype, designated here Cact. Succ. J. (Los Angeles): 165. 1949: Illustration "fig. 107. Coryphantha oklahomensis sp. nov. Photo by Jim Slack").

Coryphantha alversonii exaltissima = Coryphantha alversonii var. exaltissima Wiegand & Backeb., Cactaceae 5: 3001. 1961. Type: United States, California, ohne nähere Standortsangabe, E. F. Wiegand s. n. (lectotype, designated here Cactaceae (Backeberg): 3001. 1961: Illustration "Abb. 2817. Links: Coryphantha alversonii (Coult.) Orc.; rechts: deren v. Coryphantha exaltissima Wieg & Backbg. (photo: E. F. Wiegand.)").

Coryphantha vivipara kaibabensis = Coryphantha vivipara var. kaibabensis P.C.Fisch., Cact. Succ. J. (Los Angeles) 51: 287. 1979. Escobaria vivipara var. kaibabensis (P.C.Fisch.) N.P.Taylor, Kakteen And. Sukk. 34: 139. 1983. Type. United States, Arizona, P. C. Fischer 4094. (holotype: UC).

Coryphantha vivipara buoflama = Coryphantha vivipara var. buoflama P.C.Fisch., Cact. Succ. J. (Los Angeles) 52: 28. 1980. Escobaria vivipara var. buoflama (P.C.Fisch.) N.P.Taylor, Kakteen And. Sukk. 34: 140. 1983. Type. United States, Arizona, Yavapai County, 05 May 1979, P. C. Fischer 6582. (holotype: ARIZ; isotype: ASU [image 018464!]).

Coryphantha vivipara bisbeeana sonorensis = Coryphantha vivipara var. bisbeeana f. sonorensis P.C.Fisch., Cact. Succ. J. (Los Angeles) 52: 191. 1980. Type. México, Sonora, 84 km north of Nacozari, on the road to U.S. border, 1430 m, 27 Apr 1971, P. C. Fischer 4364. (holotype: UC).