Vietoderus Fedorenko, 2023

Fedorenko, D. N., 2023, New taxa of Pterostichus (Coleoptera: Carabidae) from Vietnam, Russian Entomological Journal 32 (1), pp. 16-39 : 26

Subgenus Vietoderus Fedorenko , subg.n.

Type species: Pterostichus laevibasis Fedorenko, sp.n.

DIAGNOSIS. A monobasic subgenus of Pterostichus different from Neohaptoderus in the following points: head rather flat; mandibles smooth dorsally (vs. densely obliquely striated); reflexed lateral margin of elytra dull from microsculpture much coarser than that on elytral intervals (vs. elytra subequally and superficially microsculptured throughout); humeral tooth nearly indistinct; ti 3 without (vs. with) lateral setae; abdominal sternite VII with apical setae distant from (vs. proximate to) apical margin in both sexes; abdominal sternite VII modified into a large median tubercle (vs. not or barely modified) in male; right paramere short; aedeagus in lateral view less curved; internal sac with a hypertrophied, transverse, preapical bulb having a few vesicles (vs. rather simple, with rather small sclerotized vesicles only). Tergite VIII and sternite VIII wide (long) in female, sternite VIII glabrous apically ( Figs 58–59 ). Spermatheca differentiated.

NAME. An abbreviated combination of Vietnam, the country from where the type species originates, and Neohaptoderus , the most similar subgenus of Pterostichus .

COMMENTS. In the East Asia, the new subgenus is similar to Neohaptoderus and Rhagadus Motschulsky, 1866 , in having the apically beaded prosternal process, certainly plesiomorphic and rather rare character within Pterostichus . The pronotum bisulcate on each side, the elytra without parascutellar striole in some species, and the interval 3 with two discal setae, d1 and d2, are characters the new subgenus shares with Rhagadus . However, species of the latter subgenus have the body slenderer, the basolateral foveae punctate, the metepisterna long, and aedeagus peculiar, with right paramere long.

On the other hand, the only known species is more similar to some representatives of the subgenus Vietosteropus in appearance, with which it also shares the elytra with coarsely microsculptured reflexed lateral margins, the mandibles smooth dorsally, ti 3 without lateral setae, and the sternite VIII glabrous apically in female.

Figures

Figs 52–62. Abdominal urites VIII–IX in female: 52–53, 61 — Pterostichus austrinus sp.n.; 54–55 — P. glabellus sp.n.; 56–57 — P. unisetosus sp.n.; 58–60 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — tergite VIII; 53, 55, 57, 59 — sternite VIII; 60 — urite IX and reproductive tract, dorsal (a) and ventral (b) aspects; 61–62 — tergite IX, ventral aspect; bc — bursa copulatrix; gsc — gonosubcoxite; LT — laterotergite; ov — common oviduct; sc — seminal canal. Scale bars 1 mm. Рис. 52–62. Уриты VIII–IX брюшка самки: 52–53, 61 — Pterostichus austrinus sp.n.; 54–55 — P. glabellus sp.n.; 56–57 — P. unisetosus sp.n.; 58–60 — P. laevibasis sp.n.; 62 — P. batdaiensis sp.n.; 52, 54, 56, 58 — тергит VIII; 53, 55, 57, 59 — стернит VIII; 60 — урит IX и репродуктивный тракт, дорЗально (a) и вентрально (b); 61–62 — тергит IX, вентрально; bc — копулЯтивнаЯ сумка; gsc — гоносубкоксит; LT — латеротергит; ov — непарный Яйцевод; sc — семепровод. Масштаб 1 мм.