Ptygmatis? tornata (Quenstedt, 1852)

Gruendel, Joachim, Keupp, Helmut, Lang, Fritz & Nuetzel, Alexander, 2022, Late Jurassic (Upper Kimmeridgian) Heterobranchia (Gastropoda) of the coral-facies of Saal near Kelheim and the viciniy of Nattheim (Germany), Zitteliana 96, pp. 179-221 : 179

Ptygmatis? tornata (Quenstedt, 1852)

Morphotype 1 Plate 10: figs 1-10

?*1830-1833 - Nerinea terebra Schübler - Zieten: 48, pl. 36, fig. 2.

?*1836 - Nerinea incavata sp. nov. - Bronn: 553, pl. 6., fig. 22.

1836 - Nerinea terebra Schübl. - Bronn: 557.

v*1852 - Nerinea tornata - Quenstedt: 429, pl. 34, fig. 36.

1852 - Nerinea constricta - Quenstedt: pl. 34, fig. 32.

1858 - Nerinea tornata - Quenstedt: 757, pl. 94, figs 12, 13.

v 1881-1884 - Nerinea tornata - Quenstedt: 527, pl. 205, figs 67, 68.

v1881-1884 - Nerinea cochlearis - Quenstedt: 556, pl. 207, figs 14, 15.

1901 - Nerinea tornata Quenst. - Geiger: 295.

1997 - Nerinella tornata (Quenstedt, 1852) - Hägele: 132, fig. p. 132 lower left.

1997 - Nerinella partschi (Peters, 1855) - Hägele: 132, pl. 13, fig. 2; fig. p. 132 upper left.

2017 - Nerineoidea Nr. 3 - Gründel: 33, pl. 13E.

Lectotype designation.

Quenstedt (1852) based Nerinea tornata on a specimen from Nattheim ( Quenstedt 1852, pl. 34, fig. 36). This illustration is somewhat schematic. It is probably the same specimen that was illustrated by him elsewhere ( Quenstedt 1881-1884, pl. 205, fig. 67) and that is also illustrated herein (Plate 10 : figs 8, 9). This specimen is herewith designated as the lectotype.

Material.

Thirty-one specimens representing the typical form from Saal (collection Lang) of which five specimens are illustrated (SNSB-BSPG 2021 XV 32-36) and seven specimens representing the morphotype 2 from Saal: five specimens collection Lang of which two are illustrated (SNSB-BSPG 2021 XV 37, 38), one specimen collection Keupp (SNSB-BSPG 2021 XV 39), one specimen collection Neubauer); three specimens from Nattheim: Nerinea tornata , Tübingen (collection Quenstedt); two fragments from Nattheim: Nerinea cochlearis , Tübingen (collection Quenstedt).

Description.

The shell is very slender; a fragmentary specimen (incomplete ad- and abapically) is 62 mm high; the shell illustrated in Plate 10 : fig. 6 is ca. 50 mm high. If the assignment of the specimen illustrated in Plate 10 : fig. 1 is correct, then the juvenile shell is very long, slender, and consists of numerous whorls. The whorls are generally wider than high but the height/width ratio varies. The whorls increase only slowly in width. The whorl face is concave. The only recognizable ornament is a subsutural bulge forming the whorl periphery. The subsutural bulge forms a sharp crest and ramp in well-preserved specimens. The whorls have a subsutural bulge bordered by the adapical suture. The base is flat, smooth and joins the whorl face at an angular edge. The base has a narrow umbilicus. The growth lines are generally opisthocline and curve backward strongly immediately below the adapical suture. The damaged aperture has a rhomboid outline and an oblique siphonal canal, two columellar plaits, one parietal, and one palatal plait.

Figures

Plate 10. Ptygmatis? tornata (Quenstedt, 1852). (1 - 10) Ptygmatis? tornata (Quenstedt, 1852) morphotype 1. (1) SNSB-BSPG 2021 XV 32; Saal (collection Lang); lateral view, height 14 mm. (2) SNSB-BSPG 2021 XV 33, Saal (collection Lang), lateral view, height 65 mm (original to Gruendel 2017, fig. 13 E). (3 - 4) SNSB-BSPG 2021 XV 34, Saal (collection Lang); (3) lateral view, height 20 mm; (4) base, width 7 mm). (5) SNSB-BSPG 2021 XV 35, Saal (collection Lang), lateral view, height 19 mm. (6 - 7) SNSB-BSPG 2021 XV 36, Saal (collection Lang); (6) lateral view, height ca. 50 mm; (7) aperture, width 15 mm. (8 - 9) Lectotype; University Tuebingen (collection Quenstedt), Nattheim, shell in lateral views, height ca. 30 mm (original Quenstedt 1881 - 1884, pl. 205, fig. 67). (10) University Tuebingen (collection Quenstedt), Nattheim, lateral view, height ca. 45 mm (original Quenstedt 1881 - 1884, pl. 206, fig. 68). (11 - 12) Ptygmatis? tornata (Quenstedt, 1852) morphotype 2; (11) SNSB-BSPG 2021 XV 37, Saal (collection Lang), lateral view, height 29 mm. (12) morphotype 2, SNSB-BSPG 2021 XV 38, Saal (collection Lang), lateral view, height 42 mm (original to Gruendel 2017, fig. 13 D).