Stryphnodendron glandulosum (Forero) Scalon, 2022

Scalon, Viviane Renata, Paula-Souza, Juliana De, Lima, Alexandre Gibau De & Souza, Vinicius Castro, 2022, A synopsis of the genus Stryphnodendron (Fabaceae, Caesalpinioideae, mimosoid clade), Phytotaxa 544 (3), pp. 227-279 : 245-246

12. Stryphnodendron glandulosum (Forero) Scalon , comb. et stat. nov.

≡ Stryphnodendron guianense (Aubl.) Benth. subsp. glandulosum Forero (1972: 145) .—Type: BRAZIL. Pará, “Museu Paraense, Cult. et Peruvia orientalis (Rio Huallaga J. Huber anno 1898)”, September 1936, A. Ducke 274 (holotype NY 00003368!, isotypes K!, R!, US!)

Trees 8–20 m tall, rhytidome smooth, gray to dark-brown. Leaves with (7–)9–12 pairs of pinnae; (7–)9–18 pairs of leaflets; petiolar nectary 1, verruciform; leaflets (5–)7-13 × (2.5–) 3-5 mm, usually oblong, distal leaflets usually obovate, proximal leaflets sometimes elliptic, chartaceous, discolor, upper surface puberule to pubescent, lower surface densely puberulent to pubescent, bilateral tuft of trichomes on the lower surface present, usually prolonged up to the middle part of the blade. Inflorescences of simple thyrsi; cymulae of geminate spikes; spikes 10–11 cm long. Flowers monoclinous, corolla tube sparsely pubescent, lobes yellow-pubescent, yellowish. Fruit a nucoid legume, slightly curved, subturgid, seeds slightly prominent, valves subcoriaceous, inconspicuously nerved.

Vernacular names: — Pashaco, Pashaca ( Peru); Taco-macho, Taco-blanco, Taco-macino ( Bolivia); Fava ( Brazil).

Remarks: — Stryphnodendron guianense var. glandulosum has always been considered an atypical taxon within the species, and our analysis corroborates that the diagnostic features of this variety are too distrepant from S. guianense var. guianense with respect to number and position of nectaries, indument of flowers and foliolules, as well as by their allopatric distribution. For this reason, S. guianense var. glandulosum is elevated as a distinct species.

Distribution and habitat: —The species is distributed only in Western Amazon (Acre state in Brazil, northern Bolivia, western Peru and southern Colombia), in areas of tropical forest of terra firme, sometimes near watercourses in seasonally flooded areas. The few records of this species outside these areas are cultivated trees in the Emílio Göeldi Museum (Belém, Pará state in Brazil), and do not correspond to the natural distribution of S. glandulosum .

Conservation:—Least Concern (LC) [AOO= 52.000 km 2, EOO= 311,438.075 km 2].

Phenology: — Stryphnodendron glandulosum was collected with flowers in September, November, January and February and with fruits from May to September.

Selected specimens examined: — BOLIVIA. Ballivian , Triunfo, Rio Matos, 29 August 1976, fr., E. Meneces & W. Terceros 07 ( INPA) . PERU. Manu , Parque Nacional Manu , Rio Manu, Cocha Cashu Station, 17 September 1986, fl., R.B. Foster 11402 (F) .