Spelaeodiscus albanicus albanicus (A. J. Wagner, 1914)

Pall-Gergely, Barna, Deli, Tamas, Eross, Zoltan Peter, Reischuetz, Peter L., Reischuetz, Alexander & Feher, Zoltan, 2018, Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae), ZooKeys 769, pp. 13-48 : 13

Spelaeodiscus albanicus albanicus (A. J. Wagner, 1914) Figure 3

Aspasita albanica A. J. Wagner, 1914 in Sturany & Wagner 1914: 67, plate 2, figs 10a-c.

Spelaeodiscus albanicus - Bole 1965: 354, Plate 76, fig. B.

Spelaeodiscus (Spelaeodiscus) albanicus - Gittenberger 1969: 294, fig. 2.

Spel aeodiscus albanicus - Pilsbry 1926: 184, plate 22, figs 12-14.

Spelaeodiscus albanicus - Welter-Schultes 2012: 213 (partim: locality data from Montenegro are of unknown origin).

Spelaeodiscus albanicus - Reischütz et al. 2013: 62, fig. 3.

Type material.

Kiri-Brücke nächst Mesi b. Skutari, Albanien (im Genist), leg. Sturany, 27.04.1905, NHMW 43385 (lectotype, hereby selected, SW: 3.7 mm, SH: 2 mm, Fig. 3A-F ); Drinasca-Ufer b. Skutari (angeschwemmt), leg. Sturany, 03.05.1905, NHMW 112351 (1 corroded, juvenile paralectotype).

Other material.

Vrelo Pronisicut (or Pronifkut), coll. Edlauer ex coll. Kuščer, NHMW 48236/1 shell ( “photo”); Pronisicut, coll. Edlauer ex coll. Kuščer, NHMW48336/2 shells; Albania, Shkodër district, Drisht, right bank of Kir river opposite to the fortress hill, 90 m a.s.l. (roadside limestone rocks), 42°7.824'N, 19°36.540'E, (site code: 2016/32), leg. Z.P. Erőss, Z. Fehér, J. Grego & M. Szekeres, 28.06.2016, HNHM 103195/1 (photographed shell, Fig. 3G-L ), NHMW 112356/ 1 adult + 5 juvenile shells ; Albania, Shkodër district, 1 km NE of Ura e Shtrenjtë, 160 m a.s.l., (roadside limestone rocks), 42°9.180'N, 19°40.000'E (site code: 2016/33), leg. Z.P. Erőss, Z. Fehér, J. Grego & M. Szekeres, 28.06.2016, HNHM 103196/1 (photographed shell, Fig. 3M-R ), NHMW 112357/ 9 juvenile /broken shells ; Albania, Shkodër district, 4 km SW of Prekal , 170 m a.s.l., (roadside limestone rocks), 42°9.936'N, 19°41.334'E (site code: 2016/35), leg. Z.P. Erőss, Z. Fehér, J. Grego & M. Szekeres, 28.06.2016, JG/ 1 adult + 1 juvenile shell ; Albania, Ura e Mesit, debris of the Kir river , 50 m a.s.l., 42°6.870'N, 19°34.498'E, leg. A. Reischütz, N. Reischütz & P. L. Reischütz, Apr. 2012, REI/1 ; Albania, rocks southwest of Zusi , southwest of Skoder, 13 m a.s.l., 42°2.316'N, 19°28.866'E, leg. A. Reischütz, N. Reischütz & P. L. Reischütz, May 2015, NHMW 112358/1 (photographed shell, Fig. 3S-X ), REI/5 ; Albania, Felsspalten above Drisht at Mes , 115 m a.s.l., 42°7.735'N, 19°36.823'E, leg. A. Reischütz, N. Reischütz & P. L. Reischütz, Jul. 2010, REI/ 2 juvenile shells ; Albania, ruins 3 km above Drisht at Mes , 195 m a.s.l., 42°7.468'N, 19°36.846'E, leg. A. Reischütz, N. Reischütz & P. L. Reischütz, Jul. 2010, REI/3 ; Albania, Periferi Shkodër, ca. 18 km upstream from dam at Koman, a left side-valley of Liqeni i Komanit , 170 m a.s.l., limestone rocks, debris, 42°13.613'N, 19°54.300'E, leg. Z.P. Erőss, Z. Fehér, A. Hunyadi & D. Murányi, 15 Apr. 2006, HNHM 102244/1 .

Diagnosis.

Protoconch matte; aperture with a rather weak palatal and two weak basal teeth.

Description.

Shell rarely flat, usually spire somewhat elevated; protoconch consists of 1.5-1.75 whorls, very finely granulated, rather matte, not glossy; teleoconch with strong, equidistant ribs that are supported by fine periostracal filaments in fresh shells; rib density variable (43-93 ribs on body whorl), usually widely spaced; between main ribs some fine wrinkles discernible; entire shell with 4.25-4.5 whorls; aperture semilunar or due to the straight basal part triangular; peristome expanded and slightly reflected, especially on the palatal, basal and umbilical areas; palatal tooth of variable strength, usually weak, although present in all adult shells, palatal region of peristome without outer incision; basal portion of peristome usually straight, slightly thickened, with two low denticles that are visible in all adult shells; umbilicus funnel-shaped, wide (although width depends on spire height).

Measurements. SW: 3.6-4.3 mm (median = 4.0 mm), SH: 1.9-2.2 mm (median = 2.0 mm), AW: 1.3-1.7 mm (median = 1.5 mm), AH: 1.3-1.6 mm (median = 1.5 mm), (n = 6; largest and smallest specimens of multiple populations measured).

Differential diagnosis.

See under S. albanicus edentatus ssp. n. Densely ribbed specimens of this subspecies might resemble large specimens of S. unidentatus . The latter species, however, usually has stronger teeth, and narrower aperture.

Variation among specimens.

Some variability was found between populations in terms of rib density, spire height, and strength of apertural teeth.

Distribution.

This subspecies is distributed in northwestern Albania. Most of the known distribution records are from the Kir River Valley. One shell found in the fluvial debris of the Drin River above the Koman Dam extends the range farther eastwards (Figure 4 ). Welter-Schultes (2012) erroneously reports this taxon from Montenegro.

Conservation status.

See under Spelaeodiscus albanicus .

Figures

Figure 3. Shells of Spelaeodiscus albanicus albanicus (A. J. Wagner, 1914). A-F lectotype (NHMW 43385) G-L Albania, Drisht, right bank of Kir river opposite to the fortress hill (HNHM 103195) M-R Albania, Shkoder district, 1 km NE of Ura e Shtrenjte (HNHM 103196) S-X Albania, rocks southwest of Zusi (NHMW 112358).

Figure 4. Distribution of Spelaeodiscus Brusina, 1886 species.