ASTROBLEPIDAE Bleeker, 1862

Published, First, 2007, Checklist of catfishes, recent and fossil (Osteichthyes: Siluriformes), and catalogue of siluriform primary types, Zootaxa 1418, pp. 1-628 : 63-68

ASTROBLEPIDAE Bleeker, 1862

Argeini Bleeker, 1862 (in Bleeker, 1862–63): 15. Type genus: Arges Valenciennes, 1840 .

Astroblepiformes Bleeker, 1862 (in Bleeker, 1862–63): 15. Type genus: Astroblepus Humboldt, 1805 .

Cyclopidae Eigenmann, 1910: 416 . Type genus: Cyclopium Swainson, 1838 .

Remarks: The name Astroblepiformes takes precedence over the simultaneously published Argeini, because it was proposed at a higher taxonomic level (subfamily vs. stirps).

Taxonomic summary: Schaefer (2003b).

1 genus, 54 species; no named fossil taxa.

ASTROBLEPUS Humboldt, 1805

Astroblepus Humboldt, 1805a: 19 . Type species: Astroblepus grixalvii Humboldt, 1805 . Type by monotypy. Gender: Masculine.

Cyclopium Swainson, 1838: 355 . Type species: Cyclopium humboldtii Swainson, 1839 . Type by subsequent monotypy. Gender: Neuter. Genus first proposed without species; species first added in Swainson (1839: 305).

Arges Valenciennes , in Cuvier & Valenciennes, 1840b: 333 (247 in Strasbourg deluxe edition). Type species: Arges sabalo Valenciennes, 1840 . Type by subsequent designation by Bleeker (1862–63: 15). Gender: Masculine.

Brontes Valenciennes , in Cuvier & Valenciennes, 1840b: 341 (254 in Strasbourg deluxe edition). Type species: Brontes prenadilla Valenciennes, 1840 . Type by monotypy. Gender: Masculine. Preoccupied by Brontes Fabricius, 1801 , in Coleoptera ; replaced by Strephon Gistel, 1848 .

Strephon Gistel, 1848 : viii. Type species: Brontes prenadilla Valenciennes, 1840 . Type by being a replacement name. Gender: Masculine. Replacement for Brontes Valenciennes, 1840 , preoccupied by Brontes Fabricius, 1801 , in Coleoptera .

Stygogenes Günther, 1864: 223 . Type species: Cyclopium humboldtii Swainson, 1839 . Type by subsequent designation of Jordan (1919b: 332). Gender : Masculine.

Astroblepus boulengeri (Regan, 1904)

Arges boulengeri Regan, 1904b: 310 . Type locality: Canelos , Ecuador. Syntypes: BMNH 1890.6.13.12–13.

Distribution: Upper Amazon River basin of Ecuador (Schaefer, 2003b).

Astroblepus brachycephalus (Günther, 1859)

Arges brachycephalus Günther, 1859: 92 . Type locality: Andes of western Ecuador. Syntypes: BMNH 1858.7.25.37–40 (4).

Distribution: Pacific draining rivers of Ecuador (Schaefer, 2003b).

Astroblepus caquetae Fowler, 1943

Astroblepus caquetae Fowler, 1943a: 246 , fig. 23–25. Type locality: Florencia , Río Ortueguasa, Colombia. Holotype: ANSP 70506 .

Distribution: Orteguaza River, upper Amazon River basin of Colombian Andes (Schaefer, 2003b).

Astroblepus chapmani (Eigenmann, 1912)

Cyclopium chapmani Eigenmann, 1912a: 13 . Type locality: Boquia , Colombia. Holotype: FMNH 56071 ; holotype illustrated in Eigenmann (1922: 51, pl. 6, fig.2).

Distribution: Andean streams of Cauca, Magdalena and San Juan River basins, Colombia (Maldonado-Ocampo et al., 2005).

Astroblepus chimborazoi (Fowler, 1915)

Cyclopium chimborazoi Fowler, 1915a: 241 , fig. 11. Type locality: Junction of Chanchan and Chiguancay rivers. Province of Chimborazo, Ecuador. Holotype: ANSP 43523 .

Distribution: Pacific draining rivers of Ecuador (Schaefer, 2003b).

Astroblepus chotae (Regan, 1904)

Arges chotae Regan, 1904b: 313 , pl. 21 (fig. 5). Type locality: Chota Valley , northern Ecuador. Holotype: BMNH 1898.11 .4.22.

Distribution: Pacific draining rivers of Colombia, Ecuador, and Peru; Magdalena and Orinoco River basins (Schaefer, 2003b).

Astroblepus cirratus (Regan, 1912)

Arges cirratus Regan, 1912b: 670 . Type locality: Río Condoto , Río San Juan, S.W. Colombia. Holotype: BMNH 1912.3 .2.7.

Distribution: San Juan River basin, Colombia (Maldonado-Ocampo et al., 2005).

Astroblepus cyclopus (Humboldt, 1805)

Pimelodus cyclopum Humboldt, 1805b: 24 , pl. 6. Type locality: Subterranean waters in Andes of Quito (basin of Río Esmeraldas), Ecuador. Type (s): Whereabouts unknown.

Cyclopium Humboldtii Swainson, 1839: 305 . Unneeded new name for Pimelodus cyclopum Humboldt, 1805 (but listed as Pimelodus cyclopium ).

Stygogenes humboldtii Günther, 1864: 223 . Type locality: Azores [in error]. Syntypes (4): BMNH 1855.6 .26.14.

Distribution: Esmeraldas River and other Pacific coastal drainages of Ecuador (Schaefer, 2003b); Andean streams of Colombia (Maldonado-Ocampo et al., 2005).

Astroblepus eigenmanni (Regan, 1904)

Arges eigenmanni Regan, 1904b: 312 , pl. 21 (fig. 3). Type locality: Cayambe and Machachi, Ecuador. Syntypes: BMNH 1892.5.14.1–8 (8).

Distribution: Pacific coastal drainages of Ecuador (Schaefer, 2003b).

Astroblepus festae (Boulenger, 1898)

Arges festae Boulenger, 1898c: 9 . Type locality: Vallée de Gualaquiza , Equateur Oriental. Syntypes: BMNH 1898.11.4.23–29; MZUT 1486 , MZUT 1497 .

Distribution: South America: Upper Marañon River of southern Ecuador (Schaefer, 2003b).

Astroblepus fissidens (Regan, 1904)

Arges fissidens Regan, 1904b: 316 . Type locality: Andes of Ecuador. Syntypes: BMNH 1887.1 . 7.2, BMNH 1889.2 .1.4397.

Distribution: Andes of Ecuador (Schaefer, 2003b).

Astroblepus formosus Fowler, 1945

Astroblepus formosus Fowler, 1945c: 10 , fig. 1–2. Type locality: Acobamba , near Tarma, at 2900 meters elevation, Río Ucayali basin, Peru, Holotype: ANSP 71647 .

Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).

Astroblepus frenatus Eigenmann, 1918

Astroblepus frenatus Eigenmann, 1918b: 676 . Type locality: Quebrada de San Joaquín , Santander, Colombia. Holotype: FMNH 58384 .

Distribution: Streams of Magdalena, Catatumbo and Orinoco river basins, Colombia (Maldonado-Ocampo et al., 2005).

Astroblepus grixalvii Humboldt, 1805

Astroblepus grixalvii Humboldt, 1805a: 19 , pl. 7. Type locality: Río Palacé , near Popayan, Colombia. Type (s): Whereabouts unknown.

Distribution: Cauca and Magdalena River basins, Colombia (Maldonado-Ocampo et al., 2005).

Astroblepus guentheri (Boulenger, 1887)

Stygogenes guentheri Boulenger, 1887b: 348 . Type locality: New Granada. Syntypes: BMNH 1880.2.26.18–25 (8).

Distribution: Andean streams of the Cauca and Magdalena River basins, Colombia (Maldonado-Ocampo et al., 2005).

Astroblepus heterodon (Regan, 1908)

Arges heterodon Regan, 1908f: 799 , pl. 49 (fig. 2, 2a, b). Type locality: Jimenez , western Colombia. Holotype: BMNH 1908.5 .29.80.

Distribution: Pacific draining rivers of Colombia (Schaefer, 2003b).

Astroblepus homodon (Regan, 1904)

Arges homodon Regan, 1904b: 309 , pl. 21 (fig. 1). Type locality: Villeta , Colombia, 3500 ft. Holotype: BMNH 1902.5 .15.27.

Distribution: Andean streams of the Cauca and Magdalena River basins, Colombia (Maldonado-Ocampo et al., 2005).

Astroblepus jurubidae Fowler, 1944

Astroblepus jurubidae Fowler, 1944b: 237 , fig. 10–13. Type locality: clear brook, tributary to Río Jurubidá [northwestern Colombia]. Holotype: ANSP 71431 .

Distribution: Pacific draining rivers of Colombia (Schaefer, 2003b).

Astroblepus labialis Pearson, 1937

Astroblepus labialis Pearson, 1937: 95 , pl. 13 (fig. 4). Type locality: Balsas , Peru. Syntypes: CAS 64617 (7).

Distribution: Marañon River basin, Peru (Schaefer, 2003b).

Astroblepus latidens Eigenmann, 1918

Astroblepus latidens Eigenmann, 1918b: 674 . Type locality: Piperel , Colombia. Holotype: FMNH 58366 ; holotype illustrated in Eigenmann (1922: 228, pl. 6, fig. 3).

Distribution: Upper Meta River basin, Colombia (Schaefer, 2003b); and Suarez River basin, Colombia (Maldonado- Ocampo et al., 2005).

Astroblepus longiceps Pearson, 1924

Astroblepus longiceps Pearson, 1924: 15 , pl. 2 (fig. 5). Type locality: Río Colorado , Lower Bopi, Bolivia. Holotype: CAS 64618.

Distribution: Upper Madeira River basin of Bolivia and Peru (Schaefer, 2003b).

Astroblepus longifilis (Steindachner, 1882)

Arges longifilis Steindachner, 1882b: 177 . Type locality: Río Huambo und de Tortora [ Peru]. Syntypes (4): at NMW. Species described in more detail and illustrated in Steindachner (1882c: 19, pl. 5, figs. 3, 3a, 3b).

Distribution: Upper Huallaga River basin, Peru (Schaefer, 2003b); Magdalena and Cauaca River basins, Colombia (Maldonado-Ocampo et al., 2005).

Astroblepus mancoi Eigenmann, 1928

Astroblepus mancoi Eigenmann , in Myers , 1928: 85. Type locality: Río Comberciato, Urubamba, Peru, 1800 feet. Syntypes: CAS 64620 (5), MCZ 30969 (11) , MCZ 30970 (3).

Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).

Astroblepus mariae (Fowler, 1919)

Cyclopium mariae Fowler, 1919a: 125 , pl. 8. Type locality: Brook near the small village of Choachi, 25 kilometers east of Bogota and at 1800 m. of altitude, Colombia. Holotype: ANSP 49368 .

Distribution: Upper Meta River basin, Colombia (Schaefer, 2003b).

Astroblepus marmoratus (Regan, 1904)

Arges marmoratus Regan, 1904b: 314 , pl. 21 (fig. 6). Type locality: St. Augustin , Andes of Colombia, 5000 ft. Syntypes: BMNH 1899.8.21.10–11 (2).

Distribution: Upper Orinoco River basin, Colombia (Schaefer, 2003b).

Astroblepus micrescens Eigenmann, 1918

Astroblepus grixalvii micrescens Eigenmann, 1918b: 677 . Type locality: Quebrada de Agua Larga , western slope of the eastern Andes, north of Bogotá, Colombia. Holotype: FMNH 58376 ; holotype illustrated in Eigenmann (1922: 54, pl. 7, fig. 4).

Distribution: Upper Orinoco River basin (Schaefer, 2003b); widely distributed in Colombian Andes (Maldonado- Ocampo et al., 2005).

Astroblepus mindoensis (Regan, 1916)

Cyclopium mindoense Regan, 1916: 80 . Type locality: Mindo , western Ecuador. Holotype: BMNH 1916.8 .1.1.

Distribution: Pacific coastal drainage of Ecuador (Schaefer, 2003b).

Astroblepus nicefori Myers, 1932

Astroblepus nicéfori Myers, 1932: 137 . Type locality: Sonsón , Antioquia, Colombia. Holotype: SU 24796 .

Distribution: Magdalena River basin (Schaefer, 2003b); Cauca River basin, Colombia (Maldonado-Ocampo et al., 2005).

Astroblepus orientalis (Boulenger, 1903)

Arges orientalis Boulenger, 1903a: 601 . Type locality: Albirregas and Milla rivers above Merida, Venezuela, 3500 m. altitude. Syntypes: BMNH 1903.6.30.81–89, USNM 133142 .

Distribution: Lake Maracaibo drainage (Schaefer, 2003b).

Astroblepus peruanus (Steindachner, 1876)

Arges peruanus Steindachner, 1876: 601 , pl. 9 (fig. 3–6). Type locality: Amable Maria , Peru. Syntypes: NMW 16974 (3).

Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).

Astroblepus phelpsi Schultz, 1944

Astroblepus phelpsi Schultz, 1944c: 283 , pl. 7 (fig. d). Type locality: Río Cobre above its mouth near La Grita, tributary of Río Quinta, latter tributary to Río La Grita, Catatumbo system, Venezuela. Holotype: USNM 121126 .

Distribution: Lake Maracaibo drainage (Schaefer, 2003b).

Astroblepus pholeter Collette, 1962

Astroblepus pholeter Collette, 1962: 311 , fig. 1–2. Type locality: Cave in Latas , 4 km north of Archidona, Napo Prov., eastern Ecuador. Holotype: USNM 196623 .

Distribution: Napo River basin, Ecuador (Schaefer, 2003b).

Astroblepus pirrensis (Meek & Hildebrand, 1913)

Cyclopium pirrense Meek & Hildebrand, 1913: 83 . Type locality: Río Cana , Cana, Panama. Holotype: FMNH 7586 .

Distribution: Cana River basin, Panama (Schaefer, 2003b).

Astroblepus praeliorum Allen, 1942

Astroblepus praeliorum Allen , in Eigenmann & Allen, 1942: 169, pl. 13 (fig. 6–7). Type locality: Tarma; Palca; Oroya; Huancachupa Creek, Húanuco, Peru. Syntypes: CAS 7509 (4) , CAS 16114 (82) , CAS 16115 (2) , CAS 47068 (10) , CAS 47069 (1).

Distribution: Upper Marañon River basin, Peru (Schaefer, 2003b).

Astroblepus prenadillus (Valenciennes, 1840)

Brontes prenadilla Valenciennes , in Cuvier & Valenciennes, 1840b: 343 (255 of the Strasbourg deluxe edition), pl. 444. Type locality: des ruisseaux qui descendent du Cotopaxi [ Ecuador], à 5000 mètres au-dessus dy niveau e l’Ocean. Syntypes (2): MNHN a-4227 (1), MNHN a-5164 (1).

Distribution: Andes of Ecuador (Schaefer, 2003b).

Astroblepus regani (Pellegrin, 1909)

Arges regani Pellegrin, 1909c: 518 . Type locality: [ Río Cariyacu , Ecuador, elev. 3100 m]. Syntypes: MNHN 1903- 0108 (1), MNHN 1903-0109 (2) , MNHN 1903-0110 (3).

Distribution: Mira River basin and Pacific coastal drainage of Ecuador (Schaefer, 2003b).

Astroblepus rengifoi Dahl, 1960

Astroblepus rengifoi Dahl, 1960a: 459 , unnumbered figure. Type locality: small brook, tributary to the Atrato River on the left side, approx. 6 ks. above the town El Carmen de Atrato, Colombia. Holotype: at ICNMHN, current whereabouts unknown.

Distribution: Atrato River basin, Colombia (Schaefer, 2003b).

Astroblepus retropinnus (Regan, 1908)

Arges retropinna Regan, 1908f: 800 , pl. 48 (fig. 2). Type locality: Jimenez , western Colombia. Syntypes: BMNH 1908.5.29.81–82 (2).

Distribution: Dagua River basin and Pacific coastal drainage of Colombia (Schaefer, 2003b).

Astroblepus riberae Cardona & Guerao, 1994

Astroblepus riberae Cardona & Guerao, 1994: 21 , fig. Type locality: Cueva de Ninabamba , Peru, 06.33ºS, 78.51ºW. Holotype: UAB unnumbered.

Distribution: Reque River basin and Pacific coastal drainage, Peru (Schaefer, 2003b).

Astroblepus rosei Eigenmann, 1922

Astroblepus rosei Eigenmann, 1922b: 55 . Type locality: Río Jequetepeque , Llallán, Peru. Syntypes: CAS 64649 (11), FMNH 58640 (1).

Distribution: Pacific coastal drainages of Cajamarca, Peru (Schaefer, 2003b); widely distributed in Colombian Andes (Maldonado-Ocampo et al., 2005).

Remarks: Eigenmann (1922: 55) stated that the type [= holotype] was a specimen deposited in the Indiana University collection, but eleven specimens were deposited there (now CAS 64649), and no indication was given in the publication as to which was the type. Therefore, all eleven, plus one specimen deposited in the Carnegie Museum collection (now FMNH 58640), must be considered syntypes.

Astroblepus sabalo (Valenciennes, 1840)

Arges sabalo Valenciennes , in Cuvier & Valenciennes, 1840b: 335 (249 in Strasbourg deluxe edition), pl. 445. Type locality: le haut Pérou. Holotype: MNHN a-4226.

Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).

Astroblepus santanderensis Eigenmann, 1918

Astroblepus cyclopus santanderensis Eigenmann, 1918b: 675 . Type locality: Quebrada de Guapota , Colombia. Lectotype: FMNH 58433 , designated by Henn (1928: 82).

Distribution: Magdalena River basin, Colombia (Schaefer, 2003b); Suarez River basin, Colombia (Maldonado- Ocampo et al., 2005).

Astroblepus simonsii (Regan, 1904)

Arges simonsii Regan, 1904b: 317 , pl. 21 (fig. 9). Type locality: Huaras , Peru, elev. 10700 ft. Syntypes: BMNH 1900.6.20.51–55 (5).

Distribution: Santa River basin and Pacific coastal drainages of Peru (Schaefer, 2003b).

Astroblepus stuebeli (Wandolleck, 1916)

Arges stübeli Wandolleck, 1916: 158 , fig. 1–4. Type locality: Lake Titicaca. Syntypes (40): MTD F977 , MTD F4639–87 .

Distribution: Lake Titicaca basin (Schaefer, 2003b).

Astroblepus supramollis Pearson, 1937

Astroblepus supramollis Pearson, 1937: 94 , pl. 13 (fig. 3). Type locality: Balsas , Peru. Syntypes: CAS 47012 (389), CAS 47015 (36) , CAS 47022 (31).

Distribution: Marañon River basin, Peru (Schaefer, 2003b).

Astroblepus taczanowskii (Boulenger, 1890)

Arges taczanowskii Boulenger, 1890b: 451 , pl. 41 (fig. 1). Type locality: Amable Maria, Peruvian Andes. Holotype: BMNH 1875.10.14.19.

Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).

Astroblepus theresiae (Steindachner, 1907)

Arges theresiae Steindachner, 1907c: 228 . Type locality: Hohen Anden von Cayendelet. Syntypes: NMW 48091 (10), NMW 92 (7).

Distribution: Pacific coastal drainages of Ecuador (Schaefer, 2003b).

Astroblepus trifasciatus (Eigenmann, 1912)

Cyclopium trifasciatum Eigenmann, 1912a: 14 . Type locality: Caldas, Colombia. Holotype: FMNH 56076 ; holotype illustrated in Eigenmann (1922: 52, fig. 15; pl. 6, fig. 6).

Distribution: Dagua River basin and Pacific coastal drainages of Colombia (Schaefer, 2003b); also Magdalena and Cauca River basins (Maldonado-Ocampo et al., 2005).

Astroblepus ubidiai (Pellegrin, 1931)

Cyclopium ubidiai Pellegrin, 1931b: 114 . Type locality: [ Lac St. Paul, near Otavalo, Imbabura Prov.,] Equateur, elev. 2600 m. Syntypes: MNHN 1930-0172 (1), MNHN 1930-0173 (1).

Distribution: Mira River basin and Pacific coastal drainages of Ecuador (Schaefer, 2003b; Vélez-Espino, 2003).

Astroblepus unifasciatus (Eigenmann, 1912)

Cyclopium unifasciatum Eigenmann, 1912a: 15 . Type locality: Caldas, Colombia. Holotype: FMNH 56079 ; holotype illustrated in Eigenmann (1922: 52, pl. 7, fig. 2).

Distribution: Dagua River basin and Pacific coastal drainages of Colombia (Schaefer, 2003b); also Magdalena and Cauca River basins (Maldonado-Ocampo et al., 2005).

Astroblepus vaillanti (Regan, 1904)

Arges vaillanti Regan, 1904b: 312 . Type locality: Huamani Peak , Ecuador, elev. 10700 ft. Syntypes: BMNH 1897.2.16.2–4 (3).

Distribution: Pacific coastal drainages of Ecuador (Schaefer, 2003b).

Astroblepus vanceae (Eigenmann, 1913)

Cyclopium vanceae Eigenmann, 1913: 421 , pl. 32 (fig. 2). Type locality: Small stream in highlands, southeast of Tarma, Peru. Holotype: FMNH 56066 .

Distribution: Upper Ucayali River basin, Peru (Schaefer, 2003b).

Astroblepus ventralis (Eigenmann, 1912)

Cyclopium ventrale Eigenmann, 1912a: 15 . Type locality: Caldas, Colombia. Holotype: FMNH 56074 .

Distribution: Dagua River basin and Pacific coastal drainage of Colombia (Schaefer, 2003b).

Astroblepus whymperi (Boulenger, 1890)

Arges whymperi Boulenger, 1890b: 451 , pl. 41 (fig. 2). Type locality: Milligalli , Andes of Ecuador. Holotype: BMNH 1887.1 .7.1.

Distribution: Esmeraldas River basin and other Pacific coastal drainages of Ecuador (Schaefer, 2003b).