AMPHILIIDAE Regan, 1911

Published, First, 2007, Checklist of catfishes, recent and fossil (Osteichthyes: Siluriformes), and catalogue of siluriform primary types, Zootaxa 1418, pp. 1-628 : 20-29

AMPHILIIDAE Regan, 1911

Amphiliidae Regan, 1911: 565 . Type genus: Amphilius Günther, 1864 .

Doumeinae Regan, 1911: 554 . Type genus: Doumea Sauvage, 1879 .

Leptoglaninae [sic, Leptoglanidinae ] Roberts, 2003: 87. Type genus: Leptoglanis Boulenger, 1902 .

Remarks: The name Amphiliidae takes precedence over the simultaneously-established Doumeinae by virtue of the higher rank initially given to the former, following Art. 24 of the Code. Roberts (2003) and Diogo (2003b) independently proposed the new subfamily name Leptoglanidinae (incorrectly formed as Leptoglaninae by Roberts) for groups of nearly identical membership. Roberts’s name was published earlier (March 14, vs. May [31]) and therefore has priority. He et al. (1999) questioned the monophyly of the Amphiliidae as used herein and, instead, restricted the name to include only Amphilius and Paramphilus and included all other taxa in the Doumeidae.

Revision: Roberts (2003, Leptoglanidinae ).

Phylogeny: Harry (1953); He et al. (1999); Diogo (2003b).

Taxonomic summary: Skelton & Teugels (1986); Skelton (1992, west Africa).

Key to genera: Poll & Gosse (1994).

12 genera, 66 species; no named fossil taxa.

AMPHILIUS Günther, 1864

Amphilius Günther, 1864: 115 , 134. Type species: Pimelodus platychir Günther, 1864 . Type by monotypy. Originally proposed as a section of Pimelodus . Gender: Masculine.

Anoplopterus Pfeffer, 1889: 15 . Type species: Anoplopterus uranoscopus Pfeffer, 1889 . Type by monotypy. Gender: Masculine.

Chimarrhoglanis Vaillant, 1897: 81 . Type species: Chimarrhoglanis leroyi Vaillant, 1897 . Type by monotypy. Gen- der: Masculine.

Review: Skelton (1984, east and southern Africa).

Amphilius atesuensis Boulenger, 1904

Amphilius atesuensis Boulenger, 1904a: 17 . Type locality: Atesu River , Gold Coast. Syntypes (4): BMNH 1903.4.24.74–77 (4).

Amphilius pictus Nichols & La Monte, 1933a: 1 , fig. 1. Type locality: Stream near Kaleata, Liberia. Holotype: AMNH 12313 .

Distribution: Gbin River, Guinea, and Pra River, from Sierra Leone to Ghana (Skelton & Teugels, 1986).

Amphilius baudoni Pellegrin, 1928

Amphilius Baudoni Pellegrin, 1928c: 453 , unnumbered figure. Type locality: rivière Passa (affluent de l’Ogoôué), Gabon. Syntypes (3): MNHN 1928-0224 (1) , MNHN 1928-0225 (1) , MNHN 1928-0226 (1) . MRAC 20649 listed as a syntype in Eschmeyer et al. (1998), but not included in original description.

Amphilius baudoni uniformis Pellegrin, 1930: 200 . Type locality: Riv. Koumbi (Ht. Ogôoué) ... Sangha. Syntypes (6): MNHN 1929-0238 (1), MNHN 1929-0239 (5). Originally as Amphilius baudoni var. uniformis .

Distribution: Ogowe River system and Sangha River, Congo basin (Skelton & Teugels, 1986).

Amphilius brevis Boulenger, 1902

Amphilius brevis Boulenger, 1902 g: 268, pl. 29 (fig. 3). Type locality: Lindi River , which flows into the Congo at the Stanley Falls. Syntypes (2): BMNH 1902.4 . 14.21 (1), IRSNB 11 (1).

Amphilius nigricaudatus Pellegrin, 1909a: 67 . Type locality: Ngomo (Ogôoué). Syntypes (3): MNHN 1909-0014 (1), MNHN 1909-0015 (1), MNHN 1909-0016 (1). Also described as new in Pellegrin (1909d: 47).

Amphilius nigricaudatus multipunctata Pellegrin, 1909a: 67 . Type locality: Ngomo (Ogôoué). Holotype: MNHN 1909-0013. Originally as Amphilius nigricaudatus var. multipunctata . Also described as new in Pellegrin (1909d: 48, fig. 2).

Distribution: Congo River system and Leome, Kouliou, and Ogowe rivers (Skelton & Teugels, 1986).

Amphilius cryptobullatus Skelton, 1986

Amphilius cryptobullatus Skelton, 1986: 279 , figs. 15–23. Type locality: Luongo River , Upper Zaire system, above and at Route 74 crossing, 10º11'S, 29º43'E, Zambia. Holotype: ROM 46800.

Distribution: Luongo River, upper Congo River basin, Zambia (Skelton, 1986).

Amphilius jacksonii Boulenger, 1912

Amphilius jacksonii Boulenger, 1912a: 602 . Type locality: the Hima River, eastern foot-hills of Ruwenzori, 3500 feet, flowing into Lake George (Ruisamba), Nile System. Holotype: BMNH 1912.10.15.47; illustrated in Boulenger (1916: 307, fig. 182), and Seegers (1996: 191, fig. 133).

Distribution: Widely distributed in central and eastern Africa (Seegers, 1996).

Remarks: Species redescribed in Walsh et al. (2000).

Amphilius kakrimensis Teugels, Skelton & Lévêque, 1987

Amphilius kakrimensis Teugels, Skelton & Lévêque, 1987: 94 , figs. 1–2. Type locality: a small tributary of the Kakrima River , Konkoure basin, at Kasseri (10º16'N, 12º28'W), Guinea. Holotype: MNHN 1986-0599 .

Distribution: Konkoure River basin, Guinea (Teugels et al., 1987).

Amphilius kivuensis Pellegrin, 1933

Amphilius kivuensis Pellegrin, 1933b: 172 . Type locality: Boulira, Kitembo (ouest du Kivu), ... Kilembo (ouest du Kivu, altitude 2000 mètres). Syntypes (8): MNHN 1932-0038 (5) , MNHN 1933-0009 (1) , MNHN 1933-0010 (1) , MNHN 1933-0011 (1).

Distribution: Rivers to west of Lake Kivu; Ruzizi River and rivers entering Lake Tanganyika (Skelton & Teugels, 1986).

Amphilius lamani Lönnberg & Rendahl, 1920

Amphilius lamani Lönnberg & Rendahl, 1920: 171 , figs. 2–3. Type locality: Kingoyi , Lower Congo. Syntypes: NRM 9910 (2).

Distribution: Lower Congo; known only from type locality (Skelton & Teugels, 1986).

Amphilius lampei Pietschmann, 1913

Amphilius lampei Pietschmann, 1913b: 190 , pl. 2 (fig. 1). Type locality: Harrar , Abessynien [ Ethiopia]. Holotype: at Weisbaden Museums.

Distribution: Webi Shebeli tributaries, near Harrar, Ethiopia (Skelton & Teugels, 1986).

Amphilius laticaudatus Skelton, 1984

Amphilius laticaudatus Skelton, 1984: 61 , fig. Type locality: Buzi River, at bridge on Inchope to Lourenço Marques road (19º55'S, 34º15'E). Holotype: AMG P5815 .

Distribution: Buzi River system, Moçambique (Skelton, 1993).

Amphilius lentiginosus Trewavas, 1936

Amphilius lentiginosus Trewavas, 1936: 71 , pl. 2 (figs. 10–11). Type locality: a brook at Cuito , Mt. Moco , Luimbale, Cuvo River system, Angola. Syntypes: BMNH 1935.3.20.189–190 (2).

Distribution: Cuvo River system, Luachimo River and Tchimenji River, Angola (Skelton & Teugels, 1986).

Amphilius longirostris (Boulenger, 1901)

Anoplopterus longirostris Boulenger, 1901f: 447 . Type locality: Hills in the Bulu country, some 75 miles in the interior from Batanga, at an elevation of between 1500–2000 feet, Cameroon. Holotype: BMNH 1901.11 .12.133.

Distribution: Cameroon and Equatorial Guinea, also tributaries of Ngoko, Congo River system, in Cameroon (Skelton & Teugels, 1986).

Remarks: Specimen listed as holotype of Anoplopterus longirostris is entered in BMNH register with locality only as Efulen, and Boulenger (1911: 260) indicated that the locality stated in original description was near Efulen.

Amphilius maesii Boulenger, 1919

Amphilius Maesii Boulenger, 1919: 160 . Type locality: Oshwe [ Congo River system]. Syntypes (7): BMNH 1919.8.28.14–16 (3), MRAC 3180-83 (4).

Distribution: Congo River basin (Skelton & Teugels, 1986).

Amphilius natalensis Boulenger, 1917

Amphilius natalensis Boulenger, 1917a: 432 . Type locality: Krantz Kloof, Natal. Syntypes (2): BMNH 1917.7 .21.1 (1).

Distribution: Ruo River, Malawi; Zambezi system of Zimbabwe, south to Umkomaas River (Natal, South Africa) (Skelton, 1993).

Remarks: Redescribed in Barnard (1943).

Amphilius opisthophthalmus Boulenger, 1919

Amphilius opisthophthalmus Boulenger, 1919: 159 . Type locality: Oshwe, Lac Léopold II. Syntypes (7): BMNH 1919.8.28.17–18 (2), MRAC 3191-94 (4).

Distribution: Oshwe on the Lukenie River (Lake Mai-Ndombe) (Skelton & Teugels, 1986).

Amphilius platychir (Günther, 1864)

Pimelodus platychir Günther, 1864: 134 . Type locality: Sierra Leone. Syntypes (4): BMNH 2001.12.5.574–577 (4), MNHN 1898-0121 (1).

Amphilius grammatophorus Pellegrin, 1913a: 237 , fig. 1. Type locality: Rivière Kikoulo, Guinée française. Syntypes: MNHN 1913-0231 (1) , MNHN 1913-0232 (1) , MNHN 1913-0233 (1) , MNHN 1913-0234 (1) , MNHN 1913-0235 (1).

Amphilius grammatophorus brevipinna Pellegrin, 1935: 465 . Type locality: Banamanan (cercle de Kissidougou), ... Friguiagbé (près Kindia); Guinée française. Syntypes (2): MNHN 1935-0194 (1), MNHN 1935-0195 (1). Orig- inally as Amphilius grammatophorus var. brevipinna . nov. var.

Amphilius grammatophorus inaequalis Pellegrin, 1935: 464 . Type locality: Friguiagbé (près Kindia) Guinée française. Syntypes (6): MNHN 1935-0191 (1), MNHN 1935-0192 (1), MNHN 1935-0193 (4). Originally as Amphilius grammatophorus var. inaequalis nov.var.

Amphilius grammatophorus marmoratus Daget, 1963: 584 . Type locality: Kana, alt. 600 m, Mont Nimba. Holotype: MNHN 1960-0119 .

Distribution: Sierra Leone, Guinea, Côte d’Ivoire (Skelton & Teugels, 1986).

Amphilius pulcher Pellegrin, 1929

Amphilius pulcher Pellegrin, 1929c: 640 , fig. 1. Type locality: rivière Louessé (fleuve Kouilou) [ Gabon]. Syntypes (3): MNHN 1929-0224 (3). MRAC 20650 listed as a syntype in Eschmeyer et al. (1998), but not listed in description.

Amphilius pulcher ephippiata Pellegrin, 1930: 200 . Type locality: Sebé (haut Ogoôué). Syntypes: MNHN 1929- 0237 (9). Originally as Amphilius pulcher var. ephippiata nov. var.

Distribution: Ogowe and Nyanga River systems, Gabon (Skelton & Teugels, 1986).

Amphilius rheophilus Daget, 1959

Amphilius rheophilus Daget, 1959: 692 , fig. 3. Type locality: Gambie à Mpantié (parc national du Niokolo-Koba); Gambie à Banharé. Syntypes (3): MNHN 1959-0147 (2), MNHN 1960-0486 (1).

Distribution: Gambia River system (Skelton & Teugels, 1986).

Amphilius uranoscopus (Pfeffer, 1889)

Anoplopterus uranoscopus Pfeffer, 1889: 16 ; illustrated in Pfeffer (1896: 31, fig. 14). Type locality: Bad bei Ushonda (Ungúu), ... Bäche bei Mhonda. Lectotype: ZMH 11944, selected by, and illustrated in, Seegers (1996: 192, fig. 137).

Chimarrhoglanis Leroyi Vaillant, 1897: 82 . Type locality: Zanguebar, torrent de Mrogoro, par 600 mètres d’altitude. Holotype: MNHN 1897-0003 .

Amphilius grandis Boulenger, 1905c: 63 , pl. 7 (figs. 3, 3a). Type locality: Chania River of Tetse , Tana system, in cold water at an altitude of 7000 feet. Syntypes (3): BMNH 1904.12.23.50–52 (3).

Amphilius Hargeri Boulenger, 1907c: 488 . Type locality: Mlangi , British Central Africa. Holotype: BMNH 1907.10 .14.8.

Amphilius krefftii Boulenger, 1911a: 356 , fig. 276. Type locality: Usambara, in rivulet running down from Anani Hills to Sigi River. Syntypes: BMNH 1909.10.19.26–27 (2).

Amphilius oxyrhinus Boulenger, 1912c: 675 , pl. 80. Type locality: Eusso Mara , a swift mountain stream, tributary of Eusso Nyiro, British East Africa. Holotype: BMNH 1912.3 .22.120.

Amphilius brevidorsalis Pellegrin, 1919a: 399 . Type locality: Rivière Revue ( Mozambique). Holotype: MNHN 1919-0488 .

Amphilius platychir cubangoensis Pellegrin, 1936a: 56 . Type locality: Cubango près de Vila da Ponte [ Angola]. Syntypes (9): MHNG 856.86 (1), MNHN 1936-0101 (1), MNHN 1936-0102 (1), MNHN 1936-0103 (3), MRAC 138769 , NMBA 5213 (1). Originally as Amphilius platychir var. cubangoensis .

Distribution: Okavango and Zambezi basins, eastern coastal Rivers south to the Mkuze basin, Natal, and throughout east and Central Africa (Skelton, 1993); Lake Rukwa basin (Seegers, 1996).

Remarks: The name Amphilius transvaaliensis that was published in the caption to figure 1 in van der Horst, (1931: 247) fails to qualify as an available name, following the provisions in Art. 13.1.1 of the Code. In addition, there is uncertainty about whether the name was intended to be published, inasmuch as copies of this paper at CAS and CU have the figure caption crossed out and replaced with the name Amphilius grandis .

Amphilius zairensis Skelton, 1986

Amphilius zairensis Skelton, 1986: 266 , fig. 1. Type locality: River Lufu , about two km from confluence with Zaire River, near Inga hydroelectric dam, 5º31.5'S, 13º37.5'E. Holotype: MCZ 50533.

Distribution: Lufu River, Congo River basin (Skelton, 1986).

ANDERSONIA Boulenger, 1900 Andersonia Boulenger, 1900d: 528 . Type species: Andersonia leptura Boulenger, 1900 . Type by monotypy. Gender: Feminine.

Slatinia Werner, 1906: 326 . Type species: Slatinia mongallensis Werner, 1906 . Type by monotypy. Gender: Feminine.

Andersonia leptura Boulenger, 1900

Andersonia leptura Boulenger, 1900d: 529 . Type locality: a pond near Koshek , Soudan. Holotype: BMNH 1907.12 .2.2545.

Slatinia mongallensis Werner, 1906: 327 . Type locality: Bahar-el-Gebel, bei Mongalla. Holotype: NMW 79741.

Andersonia brevior Boulenger, 1918: 426 . Type locality: Shari River, Ubanghi-Shari Colony , French Equatorial Africa. Holotype: BMNH 1918.11 .12.13.

Andersonia pellegrini Boulenger, 1918: 427 . Type locality: Shari River, Ubanghi-Shari Colony , French Equatorial Africa. Holotype: BMNH 1918.11 .12.13.

Distribution: Omo, Niger and Upper Nile Rivers, Lake Chad basin (Skelton & Teugels, 1986).

Remarks: The name " Andersonia brevior sp. n. " appears in the introduction of Boulenger (1918: 426) for a species described on the next page as A. pellegrini . Harry (1953) appears to be the first to have commented on the two names for the species and, therein, he treated A. pellegrini as valid.

BELONOGLANIS Boulenger, 1902

Belonoglanis Boulenger, 1902d: 50 . Type species: Belonoglanis tenuis Boulenger, 1902 . Type by monotypy. Gen- der: Masculine.

Belonoglanis brieni Poll, 1959

Belonoglanis brieni Poll, 1959: 107 , pl. 22 (fig. 3). Type locality: Stanley Pool , Rivière N'Djili. Holotype: MRAC 118539 .

Distribution: Congo River basin (Skelton & Teugels, 1986).

Belonoglanis tenuis Boulenger, 1902

Belonoglanis tenuis Boulenger, 1902d: 50 , pl. 14 (fig. 4). Type locality: Ubangi à Banzyville. Holotype: MRAC 1289 .

Belonoglanis nudipectus Lönnberg & Rendahl, 1920: 175 , fig. 6. Type locality: Lower Congo. Holotype: NRM 16118.

Belonoglanis curvirostris Pellegrin, 1922a: 74 . Type locality: Bangui (Oubanghi). Syntypes: MNHN 1921-0428 (1), MNHN 1921-0429 (1).

Distribution: Congo River basin (Skelton & Teugels, 1986).

DOLICHAMPHILIUS Roberts, 2003

Dolichamphilius Roberts, 2003: 100 . Type species: Leptoglanis brieni Poll, 1959 . Type by original designation. Gender: Masculine.

Dolichamphilius brieni (Poll, 1959)

Leptoglanis brieni Poll, 1959: 96 , pl. 24 (fig. 2). Type locality: Stanley Pool, passe devant le refuge Jipo. Holotype: MRAC 118503 .

Distribution: Stanley Pool, Congo River (Roberts, 2003).

Dolichamphilius longiceps Roberts, 2003

Dolichamphilius longiceps Roberts, 2003: 100 . Type locality: Republic du Congo (Kinshasa): Chutes Wagenia, Kisangani. Holotype: MRAC 90-47 -P-704-708.

Distribution: Chutes Wagenia on the Lualaba River, near Kisangani (Roberts, 2003).

DOUMEA Sauvage, 1879

Doumea Sauvage, 1879: 96 . Type species: Doumea typica Sauvage, 1879 . Type by original designation. Gender: Feminine.

Doumea alula Nichols & Griscom, 1917

Doumea alula Nichols & Griscom, 1917: 722 , fig. 27. Type locality: Stanleyville , Congo. Holotype: AMNH 6531 .

Distribution: Congo River basin (Skelton & Teugels, 1986).

Doumea angolensis Boulenger, 1906

Doumea angolensis Boulenger, 1906e: 347 . Type locality: the interior of Benguella , at an altitude of 4000–5000 feet, Angola. Holotype: BMNH 1906.10 .8.12.

Distribution: Quanza River system and the interior of Benguella, Angola (Skelton & Teugels, 1986).

Doumea chappuisi Pellegrin, 1933

Doumea chappuisi Pellegrin, 1933a: 114 , fig. on p. 115. Type locality: Danané ( Côte d’Ivoire). Syntypes (8): MNHN 1932-0302 (1), NMBA 4281-87 (7).

Distribution: Guinea, Liberia and Côte d’Ivoire; Cavally, Nuon, St. Paul, and Koumba River systems (Skelton & Teugels, 1986).

Doumea thysi Skelton, 1989

Doumea thysi Skelton, 1989: 6 , fig. 3. Type locality: Cross River , 27 miles from Mamfe towards Eyomojak, Cameroon, 5º46'N, 9º03'E. Holotype: MRAC 76-32 -P-2260.

Distribution: Cross River basin, Cameroon (Skelton, 1989).

Doumea typica Sauvage, 1879

Doumea typica Sauvage, 1879: 97 . Type locality: Doumé [rivière Ogôoué]. Holotype: MNHN a-0965. Described in more detail and illustrated in Sauvage (1880c: 41, pl. 3, fig. 1).

Distribution: Congo River basin and coastal rivers from Cameroon to Congo (Skelton & Teugels, 1986).

LEPTOGLANIS Boulenger, 1902 Leptoglanis Boulenger, 1902d: 42 . Type species: Leptoglanis xenognathus Boulenger, 1902 . Type by monotypy. Gender: Masculine. Remarks: Roberts (2003) removed most species from Leptoglanis and placed them, instead, into Zairichthys. Leptoglanis xenognathus Boulenger, 1902 Leptoglanis xenognathus Boulenger, 1902d: 43 , pl. 14 (fig. 1). Type locality: l’Ubangi à Banzyville. Holotype: MRAC 1287. Distribution: Congo basin.

PARAMPHILIUS Pellegrin, 1907

Paramphilius Pellegrin, 1907a: 23 . Type species: Paramphilius trichomycteroides Pellegrin, 1907 . Type by original designation. Gender: Masculine.

Paramphilius firestonei Schultz, 1942

Paramphilius firestonei Schultz, 1942: 328 , 333, pl. 36 (fig. 3). Type locality: Bromley , Harbel [ Liberia]. Holotype: USNM 118811 .

Distribution: St. Paul, Du and Borlor Rivers, Liberia (Skelton & Teugels, 1986).

Paramphilius goodi Harry, 1953

Paramphilius goodi Harry, 1953: 203 , figs. 1–2. Type locality: Lolodorf , Bikui River , Lokunje River system, Cameroon, West Africa. Holotype: SU 16970 .

Distribution: Lokunje River basin, Cameroon (Skelton & Teugels, 1986).

Paramphilius teugelsi Skelton, 1989

Paramphilius teugelsi Skelton, 1989: 2 , fig. Type locality: ruisseau affluent de Mamou ( Little Scarcies River system), Guinea, west Africa, approximately 10º09'N, 11º50'W. Holotype: MNHN 1961-0601 .

Distribution: Little Scarcies River basin, Guinea (Skelton, 1989).

Paramphilius trichomycteroides Pellegrin, 1907

Paramphilius trichomycteroides Pellegrin, 1907a: 24 . Type locality: Ditinn (Fouta-Djalon). Holotype: MNHN 1906- 0343 .

Distribution: Senegal River headwaters, near Ditinn and Mamou, Guinea (Skelton & Teugels, 1986).

Remarks: Holotype catalog number of Paramphilius trichomycteroides incorrectly reported as MNHN 1906-0243 in Eschmeyer et al. (1998).

PHRACTURA Boulenger, 1900

Peltura Perugia, 1892: 972 . Type species: Peltura bovei Perugia, 1892 . Type by monotypy. Preoccupied by Peltura Agassiz, 1846 , in trilobites; replaced by Phractura Boulenger, 1900 . Gender: Feminine.

Phractura Boulenger, 1900d: 527 . Type species: Peltura bovei Perugia, 1892 . Type by being a replacement name. Replacement for Peltura Perugia, 1892 , preoccupied by Peltura Agassiz, 1846 . Gender: Feminine.

Paraphractura Boulenger, 1902d: 47 . Type species: Paraphractura tenuicauda Boulenger, 1902 . Type by monotypy. Gender: Feminine.

Phractura ansorgii Boulenger, 1902

Phractura ansorgii Boulenger, 1902 h: 623, pl. 37 (fig. 1). Type locality: Agberi [ Nigeria]. Holotype: BMNH 1901.12 .18.7.

Distribution: Lower Niger River (Skelton & Teugels, 1986).

Phractura bovei (Perugia, 1892)

Peltura Bovei Perugia, 1892: 972 , unnumbered figures. Type locality: Congo. Holotype: MSNG 8839 .

Distribution: Lower Congo River (Skelton & Teugels, 1986).

Phractura brevicauda Boulenger, 1911

Phractura brevicauda Boulenger, 1911a: 502 , fig. 374. Type locality: Trib. of Lobi R., S. W. of Efulen , and Kribi R ., South Cameroon; Abanga R ., Ogowe. Syntypes: BMNH 1904.7.1.119-126 (8), BMNH 1908.5.30.176–177 (2), BMNH 1908.5 .25.118 (1).

Distribution: Lower Congo River (Skelton & Teugels, 1986).

Remarks: Illustrated specimen, from the Lobi River, may not be considered a holotype, even though labeled as “type” in figure caption inasmuch as the term type is also applied in the text to the single specimen from the Abanga River.

Phractura clauseni Daget & Stauch, 1963

Phractura clauseni Daget & Stauch, 1963: 95 , fig. 2. Type locality: Haute Comoé à Samago Iri. Holotype: MNHN 1960-0148 .

Distribution: Tributaries of the Comoe and Black Volta Rivers, Upper Volta, Ogun River, Nigeria, and Upper Benoue River, Cameroon (Skelton & Teugels, 1986).

Phractura fasciata Boulenger, 1920

Phractura fasciata Boulenger, 1920a: 34 , fig. 18. Type locality: Stanley-Falls. Syntypes (4): BMNH 1919.9.10.284– 285 (2), MRAC 7241–42 (2).

Distribution: Congo River basin (Skelton & Teugels, 1986).

Phractura gladysae Pellegrin, 1931

Phractura Gladysae Pellegrin, 1931a: 209 . Type locality: Haute-Louessé. Syntypes (3): MNHN 1930-0251 (1), MNHN 1930-0252 (1) , MNHN 1930-0253 (1).

Distribution: Kouilou River system; known only from type locality (Skelton & Teugels, 1986).

Phractura intermedia Boulenger, 1911

Phractura intermedia Boulenger, 1911a: 503 , fig. 375. Type locality: Ja R., at Bitye , Libi R . affl. of Ja R ., Kribi R , and Nong R . [ South Cameroon]. Syntypes: BMNH 1903.7.28.114–116 (3), BMNH 1903.7.28.209–210 (2), BMNH 1904.7.1.165–167 (3), BMNH 1904.7 .1.213 (1), BMNH 1904.10.26.49–52 (4), BMNH 1907.5 .22.187 (1), BMNH 1909.4.29.103–105 (3), BMNH 1909.12 .3.28 (1).

Distribution: Coastal Rivers of Cameroon, from Nyong to Kouilou (Skelton & Teugels, 1986).

Phractura lindica Boulenger, 1902

Phractura lindica Boulenger, 1902 g: 268, pl. 28 (figs. 3, 3a, 3b). Type locality: Lindi River , which flows into the Congo at the Stanley Falls. Syntypes (4): BMNH 1902.4 .14.25 (1), IRSNB 12 (2 or 3), MSNM 39 (1).

Phractura lukugae Boulenger, 1920c: 402 , fig. 9. Type locality: Kabeke, a village 30 miles south of Tumbwe, on the Niemba River, near the west coast of Lake Tanganyika. Holotype: BMNH 1919.7 .24.32.

Distribution: Congo River basin (Skelton & Teugels, 1986).

Phractura longicauda Boulenger, 1903

Phractura longicauda Boulenger, 1903c: 439 . Type locality: Kribi River , southern Cameroon. Holotype: BMNH 1903.7 .28.117.

Distribution: Coastal rivers of South Cameroon and Equatorial Guinea, and Congo River basin (Skelton & Teugels, 1986).

Phractura macrura Poll, 1967

Phractura macrura Poll, 1967: 269 , fig. 129. Type locality: Caungula , riv. Cassange, (8º07'S, 18º09'E), affl. riv. Uamba, Angola. Holotype: MD 6272 .

Distribution: Uamba and Cuilo Rivers, southern tributaries of Kasai, Congo River basin, Angola (Skelton & Teugels, 1986).

Phractura scaphyrhynchura (Vaillant, 1886)

Doumea scaphyrhynchura Vaillant , in Rivière, 1886: 18. Type locality: Ogôoué à Diélé. Holotype: MNHN 1886- 0424 .

Distribution: Ogowe and Congo River basins (Skelton & Teugels, 1986).

Remarks: Pellegrin (1900: 181), lists Phractura scaphyrhynchura as a new species (nov. sp.), but cites Vaillant’s description in the text.

Phractura tenuicauda (Boulenger, 1902)

Paraphractura tenuicauda Boulenger, 1902d: 48 , pl. 14 (fig. 2). Type locality: l’Ubangi à Banzyville. Syntypes (plusieurs): BMNH 1901.12.26.48–49 (2), MRAC 1297–1299 (3).

Distribution: Congo River basin (Skelton & Teugels, 1986).

PSAMMPHILETRIA Roberts, 2003

Psammphiletria Roberts, 2003: 101 . Type species: Psammphiletria nasuta Roberts, 2003 . Type by original designation. Gender: Feminine.

Psammphiletria delicata Roberts, 2003

Psammphiletria delicata Roberts, 2003: 102 , fig. 17 e. Type locality: Stanley Pool. Holotype: MRAC 118455 .

Distribution: Stanley Pool, Congo River (Roberts, 2003).

Psammphiletria nasuta Roberts, 2003

Psammphiletria nasuta Roberts, 2003: 101 , fig. 17 a–c. Type locality: Republique Centrafricaine: Central African Republic, sandy riffle in R. Ubangui near Banda , 72 km upriver from Bangui. Holotype: CAS 92624.

Distribution: Ubangui River above Bangui (Roberts, 2003).

TETRACAMPHILIUS Roberts, 2003

Tetracamphilius Roberts, 2003: 102 . Type species: Tetracamphilius pectinatus Roberts, 2003 . Type by original designation. Gender: Masculine.

Tetracamphilius angustifrons (Boulenger, 1902)

Anoplopterus angustifrons Boulenger, 1902d: 42 , pl. 10 (fig. 4). Type locality: Banzyville [Ubangi River]. Syntypes (plusieurs): BMNH 1901.12.26.39–40 (2), MRAC 1313 (1).

Distribution: Ubangi River, Congo Basin (Skelton & Teugels, 1986).

Tetracamphilius clandestinus Roberts, 2003

Tetracamphilius clandestinus Roberts, 2003: 106 , fig. 20 b. Type locality: Republique Centrafricaine: riffles in shallow sandy area of Ubangui near Banda. 2 km upstream from Bangui. Holotype: CAS 92653.

Distribution: Ubangui River above Bangui (Roberts, 2003).

Tetracamphilius notatus (Nichols & Griscom, 1917)

Amphilius notatus Nichols & Griscom, 1917: 715 , fig. 24. Type locality: Faradje , Congo. Holotype: AMNH 6711 .

Distribution: Faradje, Uele River, Congo River basin (Skelton & Teugels, 1986).

Tetracamphilius pectinatus Roberts, 2003

Tetracamphilius pectinatus Roberts, 2003: 104 , fig. 18. Type locality: Republique du Congo (Kinshasa): Congo basin, tributary of R. Luala, 29 km N of Kibunzi (8 km S of turnoff to Kubunzi, Buozi dist. ), 45º6.5'S, 13º48'E. Holotype: CAS 92627.

Distribution: Ubangui River above Bangui; Luala River tributary; and Luala River of Southern Congo (Roberts, 2003).

TRACHYGLANIS Boulenger, 1902

Trachyglanis Boulenger, 1902d: 48 . Type species: Trachyglanis minutus Boulenger, 1902 . Type by monotypy. Gen- der: Masculine.

Trachyglanis ineac (Poll, 1954)

Phractura ineac Poll, 1954: 62 , fig. 5. Type locality: riv. Loweo, environs de Yangambi (Stanleyville). Holotype: MRAC 77460 .

Distribution: Congo River basin (Yangambi, Tohuapa) (Skelton & Teugels, 1986).

Trachyglanis intermedius Pellegrin, 1928

Trachyglanis intermedius Pellegrin, 1928b: 110 . Type locality: Kasai , Congo belge. Holotype: NMBA 3755 .

Distribution: Kasai River, Congo River basin (Skelton & Teugels, 1986).

Trachyglanis minutus Boulenger, 1902

Trachyglanis minutus Boulenger, 1902d: 49 , pl. 14 (fig. 3). Type locality: l’Ubangi à Banzyville. Syntypes (4): BMNH 1901.12.26.50–51 (2), MRAC 1291–1291 A (2).

Distribution: Congo River basin (Skelton & Teugels, 1986).

Trachyglanis sanghensis Pellegrin, 1925

Trachyglanis sanghensis Pellegrin, 1925: 103 . Type locality: Ouesso (Sangha). Holotype: MNHN 1925-0145 .

Distribution: Sanga River, Congo River basin (Skelton & Teugels, 1986).

ZAIREICHTHYS Roberts, 1967

Zaireichthys Roberts, 1967: 124 . Type species: Zaireichthys zonatus Roberts, 1967 . Type by original designation. Gender: Masculine.

Revision: Roberts (2003).

Zaireichthys brevis (Boulenger, 1915)

Leptoglanis brevis Boulenger, 1915: 169 . Type locality: Rivière Lubumbashi , à Elisabethville. Syntypes (2): BMNH 1920.5 . 26.93 (1), MRAC 12702 (1).

Distribution: Congo River basin.

Remarks: Placed by Roberts (2003: 98) in the synonymy of Zairichthys rotundiceps , but comments therein indicate only that the types are “closely similar, possibly conspecific [with Z. rotundiceps ].” Therefore, Zaireichthys brevis is retained as valid herein.

Zaireichthys camerunensis (Daget & Stauch, 1963)

Leptoglanis camerunensis Daget & Stauch, 1963: 94 , fig. 1. Type locality: Bénoué à Lakdo, Cameroun. Syntypes (7): MNHN 1962-1272 (6); one specimen lost.

Distribution: Upper Benue River, Cameroon and Upper Niger River, Guinea (Roberts, 2003).

Zaireichthys dorae (Poll, 1967)

Leptoglanis dorae Poll, 1967: 211 , fig. 95. Type locality: rapides de la Luachimo, dans des flaques d’eau résiduelle, à l’occasion de la mise à sec del la partie d’aval du barrage, Angola. Holotype: MRAC 161646 .

Distribution: Luachimo, Congo Basin, Angola (Roberts, 2003).

Zaireichthys flavomaculatus (Pellegrin, 1926)

Leptoglanis flavomaculatus Pellegrin, 1926: 204 . Type locality: Kamaiembi. Holotype: MRAC 19721 .

Distribution: Lulua River, southern Congo; known only from holotype (Roberts, 2003).

Zaireichthys heterurus Roberts, 2003

Zaireichthys heterurus Roberts, 2003: 96 . Type locality: Republic du Congo (Kinshasa): R. Avokoko, affluent rive droit fleuve Zaire, km 21 route Kisangani-Wanie Rukula. Holotype: MRAC 87-42 -P-1140.

Distribution: Eastern Congo River basin (Roberts, 2003).

Zaireichthys mandevillei (Poll, 1959)

Leptoglanis mandevillei Poll, 1959: 98 , pl. 25 (fig. 2). Type locality: Stanley Pool ( Congo R., near Kinshasa). Holotype: MRAC 118532 .

Distribution: Stanley Pool, Congo River, Ubangui and Lualaba (Roberts, 2003).

Zaireichthys rotundiceps (Hilgendorf, 1905)

Gephyroglanis rotundiceps Hilgendorf, 1905: 412 . Type locality: Im Bubu bei Irangi. Lectotype: ZMB 16392, des- ignated by, and illustrated in, Seegers (1996: 196, fig. 141).

Distribution: Cunene, Okavango and Zambezi, Pungwe, Buse and Save basins, and Lake Malawi basin (Skelton, 1993); Lake Rukwa basin (Seegers, 1996).

Zaireichthys wamiensis (Seegers, 1989)

Leptoglanis wamiensis Seegers, 1989: 285 , unnumbered figure. Type locality: Kisangata -Bach bei Mvumi, 32 kilometer SW Kidete, am Wege nach Kimamba/ Kilosa im Wami-Einzug NW Morogoro, Tanzania. Holotype: ZFMK 15899 .

Distribution: Wami River basin, Tanzania (Seegers, 1989).

Zaireichthys zonatus Roberts, 1967

Zaireichthys zonatus Roberts, 1967: 124 , figs. 3–4. Type locality: Side channel of the lower rapids of the Congo River , below Stanley Pool , at Kinsuka village, within Leopoldville city limits. Holotype: SU 64126 .

Distribution: Congo River basin (Roberts, 2003).