Europiella decolor (Uhler)

SCHUH, RANDALL T., 2004, Revision of Europiella Reuter in North America, with the Description of a New Genus (Heteroptera: Miridae: Phylinae), American Museum Novitates 3463, pp. 1-59 : 26-42

Europiella decolor (Uhler)

Figures 1 , 2 , 4 , 10

? Phytocoris pallidulus Dahlbom, 1851: 211 (n. sp.; syn. by Schuh et al., 1995: 384).

Agalliastes decolor Uhler, 1893:380 (n. sp., descr.); Schuh et al., 1995: 384 (n. syn., disc. figs.).

Agalliastes apiatus Uhler, 1895: 53 (n. sp.; syn. by Schuh et al., 1995: 384).

Agalliastes signatus Uhler, 1895: 55 (n. sp.; syn. by Schuh et al., 1995: 384).

Chlamydatus bakeri Bergroth, 1898: 35 (new name for Agalliastes signatus Uhler ).

Plagiognathus albipennis var. extrema Reuter, 1901: 187 (n. var.; syn. by Schuh et al., 1995: 384).

Chlamydatus uhlerianus Kirkaldy, 1909: 390 (unnecessary new name for Agalliastes signatus Uhler ).

Europiella decolor: Van Duzee, 1917b: 415 (n. comb., cat.); Schuh, 1995: 314 (cat.).

Psallus waldeni Knight, 1923 (n. sp.; syn. by Schuh et al., 1995: 384).

Plagiognathus (Poliopterus) litoralis Wagner, 1949: 53 (n. sp.; syn. by Schuh et al., 1995: 384).

Plagiognathus (Poliopterus) litoralis f. abrotani Wagner, 1949: 53 (n. form; syn. by Schuh et al., 1995: 384).

Plagiognathus apiatus: Kelton, 1965: 1142 (n. comb.).

Europiella nigricornis Knight, 1968: 40 (n. sp.; syn. by Schuh et al., 1995: 384).

Plagiognathus (Poliopterus) larae Kerzhner, 1978: 46 (n. sp.; syn. by Schuh et al., 1995: 384).

Europiella yampae: Kelton, 1980: 341 (diag.) (misidentification).

DIAGNOSIS: Variable in size, coloration, and degree of sexual dimorphism (figs. 1, 2); often small, but some populations approaching or exceeding size of E. artemisiae ; coloration of dorsum ranging from almost uniformly white to almost completely infuscate, specimens from much of western North America often pale, eastern North American specimens always infuscate; antennal segments 1 and 2, all femora, and entire venter moderately infuscate to black, even in specimens with pale dorsum; sexual dimorphism strong in many western North American populations, males elongate and parallel­sided, females ovoid, such dimorphism much less pronounced in eastern North American populations. Antennae and hind femora always heavily infuscate; apical vesical spines diagnostic. General appearance and structure of male genitalia (fig. 2) most similar to E. artemisiae and E. lattini , but differing from the former in having a relatively shorter antennal segment 2 in males and a shorter anterior apical vesical spine, and from the latter by having a much smaller pygophore and vesica.

REDESCRIPTION: Male: Size variable, often small, body form usually elongate, more rarely elongate ovoid, total length 2.14–3.50, width pronotum 0.69–0.96. COLORATION (figs. 1, 2): Extremely variable, dorsum ranging from almost entirely white to entirely infuscate, some specimens with dorsum generally dark except base of corium and base of cuneus pale; labium pale to nearly orange, except segment 4 black; all antennal segments black; venter of body generally infuscate to nearly black; all trochanters usually pale; fore and middle femora at most weakly infuscate, hind femora often nearly black; all tibiae of pale background coloration with black spines with black bases. SURFACE AND VESTITURE (fig. 10D): Surface and vestiture as in generic diagnosis. STRUC­ TURE (fig. 10): Most populations, especially in western North America elongate, narrowbodied, nearly parallel­sided, corial margin only weakly convex; eastern North American populations often less conspicuously elongate; antennal segment 2 only slightly longer than the width of the head; pygophore relatively small in comparison to similar appearing E. lattini . GENITALIA (fig. 3): Pygophore moderately large relative to size of abdomen; posterior apical spine of vesica short, sharply curving, hook­shaped just surpassing gonopore, anterior spine conspicuously longer and weakly decurved, structure very similar to that of E. artemisiae , also similar to E. lattini , but anterior spine much longer and entire vesica smaller than in that species.

Female: Coloration as in male. Body form much more strongly ovoid. Total length 2.45–2.72, width pronotum 0.81–0.95.

HOSTS: Breeds on a variety of herbaceous and woody Artemisia spp. Also recorded from other members of the Asteraceae , although probably not breeding. All records from non­Asteraceae are certainly sitting, rather than breeding, records.

DISTRIBUTION: Holarctic. In the Nearctic widely distributed in the West from Alaska to northern Mexico, except in wetter coastal regions. Ranging south to about 35 degrees north latitude in eastern North America.

DISCUSSION: This species appears to be truly Holarctic. Because of its great variability and the flawed diagnoses of some phyline genera, E. decolor was placed in several genera under different names ( Schuh et al., 1995). Although the extremes of variation might tempt one to recognize more than one species, examination of several thousand specimens from North America—whose genitalia appear nearly invariate, and whose second antennal segment is always relatively short—has caused me to conclude that a single species is involved, no matter what the extent of variation in sexual dimorphism, size, and coloration. The alternative to recognizing a single species is to recognize many species, most with overlapping temporal, geographic, and host distributions. Studies at the molecular level might help to clarify the underlying nature of variation and thereby further test the single­species theory.

Uhler (1895) described Agalliastes apiatus from specimens collected at Fort Collins, Manitou, and Steamboat Springs, Colorado. He indicated that the taxon also occurred in Kansas. As noted by Schuh et al. (1995), the collections of the National Museum of Natural History, Washington, D.C., contain 10 specimens associated with the A. apiatus identification. One of these, the data for which are listed below under Specimens Examined, was labelled as a cotype by H. G. Barber. I have designated it as the lectotype. Schuh et al. (1995) treated E. apiatus as a junior synonym of E. decolor . They stated that the two taxa were ‘‘clearly congeneric’’, which is true, but the statement should have read ‘‘clearly conspecific’’.

Without elaboration Schuh et al. (1995) treated Europiella nigricornis Knight, 1968 as a junior synonym of E. decolor . This action was, in fact, based on comparison of the types of these nominal species and the examination of large numbers of dissected male genitalia of E. decolor .

SPECIMENS EXAMINED: Lectotype: ♀, Am. Fork, Ut., 22.6.91, P. R. Uhler Collection USNM). Types of synonyms: Agalliastes apiatus Uhler : lectotype 3, Colo., 1691, P. R. Uhler Collection (USNM). Agalliastes signatus Uhler : Type not located. Europiella nigricornis Knight : holotype 3, MERCURY, NEVADA, 6 M, VI­17–1965, H. Knight & J. Merino (USNM). Additional specimens: CANADA: Alberta: Aden, 25 mi E of Milk River, June 28, 1956, O. Peck, 13 (CNC). Calgary, June 18, 1959, Carr, 1♀ (CNC). Calgary, July 9, 1959, Carr, 13 (CNC). Calgary, August 9, 1959, Carr, 1♀ (CNC). 32 mi W of Calgary, July 18, 1970, Oman, 63 (OSU). Canmore, August 27, 1952, A.R. Brooks, 13 (CNC). Castor, August 7, 1957, A.R. and J.E. Brooks, 1♀ (CNC). Chim, September 7, 1929, J.H. Pepper, 13 (CNC). Coal Valley, August 31, 1970, L.A. Kelton, 83 (CNC). Coronation, July 28, 1973, L.A. Kelton, 3♀ (CNC). Coults, June 15, 1952, L.A. Konotopetz, Lupinus argenteus (Fabaceae) , 33 (CNC). 3.8 km S of Drumheller on Rt 56, July 14, 1990, M. D. Schwartz, 1♀ (CNC). Drumheller, June 18, 1957, A.R. Brooks, MacNay, 2♀ (CNC). Drumheller, August 11, 1957, A.R. and J.E. Brooks, 13, 5♀ (CNC). Eisenhower Junction, August 5, 1970, L.A. Kelton, 33 (CNC). Elkwater Park, June 13, 1952, L.A. Konotopetz, 13 (CNC). Elkwater Park, July 13, 1952, L.A. Konotopetz, 23 (CNC). Elkwater Park, July 26, 1952, L.A. Konotopetz, 13, 2♀ (CNC). Empress, June 7, 1957, A.R. Brooks, MacNay, 13, 3♀ (CNC). Empress, August 22, 1957, A.R. Brooks, MacNay, 2♀ (CNC). Fisher Creek A.F.S., July 26, 1973, L.A. Kelton, 13 (CNC). Geikie Road, Jasper National Park, August 29, 1932, R.H. Handford, 13, 1♀ (CNC). Glenwood, June 19, 1938, R.W. Salt, 1♀ (CNC). High Prairie, July 16, 17, 1961, A.R. Brooks, 13, 1♀ (CNC). Irvine, June 11, 1952, L.A. Konotopetz, Artemisia cana (Asteraceae) , 143, 8♀ (CNC). Jasper, August 29, 1970, L.A. Kelton, Artemisia longifolia (Asteraceae) , 13, 25♀ (CNC). Kananaskis Highway, July 20, 1974, L.A. Kelton, 1♀ (CNC). Kananaskis Highway, August 25, 1973, L.A. Kelton, 1♀ (CNC). Just W of Kananaskis, Eyrie Gap Picnic Area, July 24, 1994, M. D. Schwartz, Artemisia frigida (Asteraceae) , 153 (CNC). Lethbridge, June 4, 1939, R.W. Salt, Medicago sp. (Fabaceae) , 1♀ (CNC). Lethbridge, June 16, 1939, R.W. Salt, Medicago sp. (Fabaceae) , 23, 3♀ (CNC). Lethbridge, August 5, 1930, J. H. Pepper, 13 (CNC). Lethbridge, September 6, 1929, J. H. Pepper, 13, 1♀ (CNC). Lethbridge, September 23, 1929, J. H. Pepper, 43, 6♀ (CNC). Manyberries, June 4, 1952, L.A. Konotopetz, 23, 1♀ (CNC). Medicine Hat, June 14, 1930, J.H. Pepper, 13 (CNC). Medicine Hat, June 2, 1952, L.A. Konotopetz, A. R. Brooks, Artemisia cana (Asteraceae) , 13, 5♀ (CNC). Nordegg, July 29, 1921, J.McDunnough, E. bakeri det. H. H. Knight, 43 (CNC). Nordegg, July 31, 1921, J.McDunnough, E. bakeri det. H. H. Knight, 3♀ (CNC). Nordegg, August 31, 1970, L.A. Kelton, 13 (CNC). Onefour, June 13, 1956, O. Peck, Agropyron junceus (Poaceae) , 1♀ (CNC). Onefour, June 14, 1952, L.A. Konotopetz, 23, 7♀ (CNC). Peace River, June 18, 1961, A.R. Brooks, 2♀ (CNC). Red Deer, June 25, 1957, A.R. Brooks, MacNay, 6♀ (CNC). Robb, August 30, 1970, L.A. Kelton, 23, 1♀ (CNC). Scandia, July 26, 1956, O. Peck, E.E. Sterns, 4♀ (CNC). Steveville, August 21, 1957, A.R. and J.E. Brooks, 5♀ (CNC). Vermillion Provincial Park, August 22, 1993, M. D. Schwartz, Artemisia biennis (Asteraceae) , 63 (CNC). Wainwright, July 27, 1957, A.R. and J.E. Brooks, 13, 2♀ (CNC). Walsh, May 28, 1952, A.R. Brooks, Artemisia cana (Asteraceae) , 13, 2♀ (CNC). Waterton Lakes National Park, July 26, 1972, L.A. Kelton, 1♀ (CNC). British Columbia: 10 mi S of Merritt, July 19, 1970, Oman, 23 (OSU). 11 km N of Liard River, July 31, 1982, G.G.E. Scudder (UBC). 13 km E of Hudson Hope, June 14, 1982, R. D. Wilkie, S. G. Cannings (UBC). 13 mi E of Hudson Hope, June 14, 1982, R. D. Wilkie and S. G. Cannings, Artemisia frigida (Asteraceae) , 93, 5♀ (UBC). 14 km E of Hudson Hope, July 26, 1986, B. A. Macdonald, S. G. Cannings (UBC). 149 Mile Lake, June 22, 1961, G.G.E. Scudder (UBC). 15 km E of Hudson Hope, June 22, 1985, Bidgemast, Cannings, and Macdonald, 43, 8♀ (UBC, CNC). 15 km E of Hudson Hope, June 22, 1985, Cannings, Macdonald, Bijdemast (UBC). 18 mi N of Lytton, June 12, 1963, G.G.E. Scudder (UBC). 2 km E of Twin Lakes, July 9, 1991, G.G.E. Scudder (UBC). 2 km E of Twin Lakes, July 9, 1991, G.G.E. Scudder, 43 (UBC). 2 km W of Greenwood, August 17, 1997, G.G.E. Scudder (UBC). 21 mi N of Kitwancool, July 31, 1979, G.G.E. Scudder (UBC). 3 km E of Tahltan River, Telegraph Creek, June 22, 1985, E. Krebs, J. J. Robinson, Artemisia frigida Willd. (Asteraceae) (UBC). 30 km SW of Chetwynd, June 22, 1985, E. Bijdemast (UBC). 5 km S of Spences Bridge, July 26, 1983, G.G.E. Scudder (UBC). 5 mi W of Toad River Lodge, August 24, 1980, S. G. Cannings (UBC). 54 km N of Atlin, Atlin Lake, July 16, 1985, E. Krebs, J. J. Robinson (UBC). 7 mi E of Oliver, Indian Reservation, July 8, 1959, L. A. Kelton ( Asteraceae ) (CNC). 8 km N of Naramata, June 12, 1992, G.G.E. Scudder (UBC). 8 km N of Osoyoos, September 11, 1995, G.G.E. Scudder (UBC). Alaska Highway, km 1204, July 30, 1982, G.G.E. Scudder (UBC). Alaska Hwy mi 392, Summit Lake, 4200 ft, July 19, 1959, R. E. Leech, 13 (CNC). Alaska Hwy, km 1204, July 30, 1982, G.G.E. Scudder, 13 (UBC). Alkali Lake, August 13, 1988, G.G.E. Scudder (UBC). Alkali Lake, August 13, 1988, G.G.E. Scudder, 33 (UBC). Anahim Lake, July 18, 1978, G.G.E. Scudder (UBC). Ashcroft, Barnes Lake, August 31, 1993, G.G.E. Scudder (UBC). Atlin, 2200 m, August 17, 1955, H. J. Huckel, 23, 1♀ (CNC). Atlin, July 27, 1979, G.G.E. Scudder (UBC). Atlin, May 2, 1955, B. A. Gibbard (CNC). Beaumont Provincial Park, Fraser Lake, July 24, 1983, G.G.E. Scudder (UBC). Blue River Bridge, km 705.6 Cassiar Highway, July 21, 1983, G.G.E. Scudder (UBC). Blue River, Stewart­Cassiar Road, August 25, 1981, C.S. Guppy (UBC). Cariboo, Long Lake, August 13, 1988, G.G.E. Scudder, 5♀ (UBC). Cariboo, White Lake , August 13, 1988, G.G.E. Scudder (UBC). Cassiar Hiway, km 705.6, Blue River Bridge, July 21, 1983, G.G.E. Scudder, 3♀ (UBC). Chopaka, January 7, 1981, S. G. Cannings, 13 (UBC). Cormier Creek, July 29, 1979, G.G.E. Scudder (UBC). Dean River, July 16, 1978, G.G.E. Scudder (UBC). Douglas Lake, September 1, 1993, G.G.E. Scudder (UBC). Douglas Lake, September 1, 1993, G.G.E. Scudder, 30♀ (UBC). Elk River, Grasmere, July 10, 1981, R. J. Cannings, 13, 5♀ (UBC). Elko, July 8, 1970, L. A. Kelton (CNC). Elko, July 8, 1970, L. A. Kelton, 1♀ (CNC). Fairmont Hot Springs, Dutch Creek, July 1, 1982, B. A. Macdonald, 13, 4♀ (UBC). Fairmont Hot Springs, Dutch Creek, July 1, 1982, G.G.E. Scudder (UBC). Fernie, July 7, 1970, L. A. Kelton (CNC). Fernie, July 7, 1970, L. A. Kelton, 83, 3♀ (CNC). Fort Nelson, August 2, 1982, G.G.E. Scudder (UBC). Fourth of July Creek, July 28, 1979, G.G.E. Scudder (UBC). Grasmere, Elk River, July 10, 1981, S. G. Cannings (UBC). Hagensborg, July 17, 1978, G.G.E. Scudder (UBC). Haynes Ecology Reserve, sandy road, August 26, 1989, D. L. Sanford (UBC). Hedley, July 29, 1978, J. A. van Reenen, 33, 2♀ (UBC). Hedley, July 29, 1978, J. A. Van Reenen, Artemisia sp. (Asteraceae) (UBC). Hedley, June 4, 1961, G.G.E. Scudder (UBC). Hedley, Nickel Plate Mine, 5800 ft, July 31, 1934, A. N. Gartrell (CNC). Highway 7 at Yukon border, July 27, 1979, G.G.E. Scudder (UBC). Hope Mountains, 4500 ft, August 22, 1932, A. N. Gartrell (CNC). Hope Mountains, 4500 ft, August 26, 1932, A. N. Gartrell (CNC). Hope Mts., 4500 ft, August 26, 1932, A. N. Gartrell, 13, 1♀ (CNC). Jaffray, July 17, 1974, L. A. Kelton (CNC). Kaleden, July 2, 1975, L. A. Kelton (CNC). Kaleden, July 2, 1975, L. A. Kelton, 1♀ (CNC). Kamloops, June 11, 1972, N. L. H. Krauss (USNM). Kamloops, June 24, 1950, B. P. Beirne ( Asteraceae ) (CNC). Kamloops, June 24, 1950, B. P. Beirne, 13, 1♀ (CNC). Kamloops, Lac du Bois, August 6, 1954, W. Downes (UBC). Kamloops, Lac du Bois, August 7, 1956, W. Downes (UBC). Kamloops, Lac du Bois, August 8, 1954, W. Downes (UBC). km 1204 Alaska Highway, July 30, 1982, G.G.E. Scudder (UBC). Ladysmith, July 13, 1959, L. A. Kelton (CNC). Lillooet, Seton Lake, June 6, 1997, G.G.E. Scudder (UBC). Lilooet, Seton Lake, June 6, 1997, G.G.E. Scudder, 123, 8♀ (UBC). Lower Post, July 20, 1983, G.G.E. Scudder (UBC). Lytton, August 2, 1931, R. H. Beamer, 23 (KU). Manning Prov. Park, Blackwall, July 23, 1970, L. A. Kelton, 23 (CNC). Manning Provincial Park, August 27, 1961, G.G.E. Scudder (UBC). Manning Provincial Park, Blackwall, August 12, 1970, L. A. Kelton (CNC). Manning Provincial Park, July 29, 1962, G.G.E. Scudder (UBC). Merritt, August 29, 1962, G.G.E. Scudder (UBC). Merritt, September 21, 1962, G.G.E. Scudder (UBC). Midway, August 10, 1970, L. A. Kelton (CNC). Midway, August 10, 1970, L. A. Kelton, 53, 4♀ (CNC). Minnie Lake, June 13, 1963, G.G.E. Scudder (UBC). Minnie Lake, June 13, 1963, G.G.E. Scudder, 15♀ (UBC). Mount Robson Provincial Park, August 27, 1970, L. A. Kelton (CNC). Mt. Kobau, 11 km NW of Osoyoos Summit, 1834 m, August 30, 1993 – August 26, 1986, M. D. Schwartz, G. E. Scudder, Artemisia tridentata (Asteraceae) , 43 (AMNH,KU). Near Haynes Ecology Reserve, July 15, 1990, G.G.E. Scudder, Artemisia tridentata (Asteraceae) (UBC). Near Osoyoos, Anarchist Mountain, 2000 ft, June 13, 1957, N. H. Anderson (CNC). Near Osoyoos, Anarchist Mountain, July 13, 1970, L. A. Kelton (CNC). Near Osoyoos, Anarchist Mountain, June 7, 1959, R. Madge (CNC). Nicola Lake, July 26, 1970, L. A. Kelton, 2♀ (CNC). Nicola, September 1, 1993, G.G.E. Scudder (UBC). Nimpo Lake, July 18, 1978, G.G.E. Scudder (UBC). Nimpo Lake, July 18, 1978, G.G.E. Scudder, 53, 5♀ (UBC). Okanangan Valley, June 1, 1996, R. G. Foottit and D. R. Gillespie ( Fabaceae ) (CNC). Oliver, 7 mi E of Indian Reservation, July 8, 1959, L. A. Kelton, 53, 20♀ (CNC). Oliver, July 19, 1970, L. A. Kelton, Artemisia (Asteraceae) (CNC). Oliver, June 27, 1959, L. A. Kelton (CNC). Oliver, Madden Lake (CNC). Oliver, May 29, 1959, L. A. Kelton (CNC). Oliver, May 29, 1959, R. E. Leech, 83, 5♀ (CNC). Oliver, Mc­ Intyre Creek, July 4, 1959, L. A. Kelton, 3♀ (CNC). Oliver, Meyer Flats, June 29, 1959, L. A. Kelton, 73, 15♀ (CNC). Oliver, Spectacle Lake, June 10, 1959, L. A. Kelton (CNC). Oliver, Vaseaux Lake, May 29, 1959, E. E. MacDougall, 53, 5♀ (CNC). Osoyoos Lake, Haynes Ecology Reserve, June 30, 1981, S. G. Cannings (UBC). Osoyoos Lake, Haynes Ecology Reserve, October 4, 1992, G.G.E. Scudder (UBC). Osoyoos Lake, Northeast end, May 19, 1980, S. G. Cannings (UBC). Osoyoos, Anarchist Mt., July 13, 1970 – June 30, 1981, L. A. Kelton, S. G. Cannings, 63 (CNC, UBC). Osoyoos, east bench, July 15, 1990, G.G.E. Scudder (UBC). Osoyoos, Mount Kobau, 1885 m, August 26, 1986, G.G.E. Scudder, Artemisia tridentata (Asteraceae) (UBC). Osoyoos, Richter Pass, June 28, 1959, L. A. Kelton, 23, 4♀ (CNC). Osoyoos, Richter Pass, June 6, 1959, R. E. Leech (CNC). Osoyoos, September 30, 1987, G.G.E. Scudder (UBC). Pavilion Lake, June 7, 1997, G.G.E. Scudder (UBC). Penticton, September 21, 1919, W. Downes ( Asteraceae ) (UBC). Penticton, west bench, June 23, 1983, G.G.E. Scudder (UBC). Penticton, west bench, June 29, 1981, S. G. Cannings (UBC). Pink Mountain, August 17, 1982, L. A. Kelton (CNC). Pink Mt., August 17, 1982, L. A. Kelton, 143, 15♀ (CNC). Prophet River, August 4, 1982, G.G.E. Scudder (UBC). Shaka Lake, Eastside Res., July 6, 1996, G.G.E. Scudder, 5♀ (UBC). Skaha Lake, Eastside Reservation, July 6, 1996, G.G.E. Scudder (UBC). Soda Creek, August 21, 1983, W. Downes (UBC). Soda Creek, July 22, 1947, G. J. Spencer (CNC). Soda Creek, July 8, 1949, R. Stace­Smith (CNC). Soda Creek, June 16, 1961, G.G.E. Scud­ der (UBC). Soda Creek, June 19, 1950 – August 22, 1947, Spencer, Stace­Smith, 33, 2♀ (CNC). Soda Creek, June 19, 1950, G. J. Spencer (CNC). Spences Bridge, September 6, 1954, W. Downes (UBC). Spilimacheen Rest Area, September 12, 1992, G.G.E. Scudder, 123, 18♀ (CNC, UBC). Stewart, July 22, 1983, G.G.E. Scudder, 23 (UBC). Strawberry Creek, August 6, 1981, C. S. Guppy, 1♀ (UBC). Summerland, July 10, 1975, L. A. Kelton, 13 (CNC). Summerland, July 12, 1975, L. A. Kelton (CNC). Summerland, July 2, 1974, L. A. Kelton, alfalfa ( Fabaceae ) (CNC). Summerland, July 6, 1975, L. A. Kelton (CNC). Summerland, Trout Creek, June 5, 1959, Kelton, Madge, Leech (CNC). Summerland, Trout Creek, June 5, 1959, R. Madge and L. A. Kelton, Artemisia tridentata (Asteraceae) , 83, 8♀ (CNC). Ta Ta Creek, June 8, 1961, G.G.E. Scudder, 53, 10♀ (UBC). Tahitan, August 10, 1986, S. G. Cannings, Artemisia frigida (Asteraceae) , 13 (UBC). Terrace, August 14, 1960, W.R. Richards (CNC). Terrace, July 10, 1960, W.R. Richards (CNC). Terrace, July 24, 1960, W. R. Richards, 123, 15♀ (CNC). Terrace, July 6, 1960, W.R. Richards (CNC). Terrace, July 7, 1960, W.R. Richards (CNC). Tetsa Br. no. 1 (CNC). Upper Swan Lake, July 25, 1987, S. G. Cannings (UBC). Vasuex L., June 27, 1981, S. G. Cannings, 13 (UBC). Vernon, June 30, 1940, R. L. Usinger, 43 (UCB). W of Kleena Kleene, McClinchey Creek, July 16, 1978, G.G.E. Scudder (UBC). Wasa, August 7, 1970, L. A. Kelton, 3♀ (CNC). West Bench Penticton, June 23, 1983, S. G. Cannings, 53 (UBC). Williams Lake, August 20, 1960, W. R. Richards, 23, 5♀ (CNC). Williams Lake, June 15, 1970, G.G.E. Scudder (UBC). Winslow Experiment Station, August 13, 1925, G. J. Spencer (UBC). Yahk, July 22, 1959, L. A. Kelton, 1♀ (CNC). Yale Co.: 7 mi W of Bridesville, July 9, 1966, W. Gagne, J. Haddock, 13 (UCB). Labrador: Goose Bay, August 23, 1948, W.E. Beckel, 1♀ (CNC). Manitoba: Angusville, July 20, 1954, A.R. Brooks, L.A. Kelton, 1♀ (CNC). Aweme, July 5, 1930, N. Criddle, Artemisia ludoviciana (Asteraceae) , 23, 2♀ (CNC). Aweme, August 18, 1922, N. Criddle, 33, 8♀ (CNC). Aweme, June 27, 1930, R.M. White , 1♀ (CNC). Aweme, June 22, 1953, A.R. Brooks, L.A. Kelton, 13, 1♀ (CNC). Aweme, August 17, 1922, N. Criddle, 1♀ (CNC). Aweme, June 9, 1921, R.M. White , 1♀ (CNC). Aweme, June 25, 1930, R.M. White , 2♀ (CNC). Aweme, August 2, 1931, R.R. Handford, 2♀ (CNC). Aweme, August 3, 1931, R.R. Handford, 1♀ (CNC). 1.0 km SE of Jct of Hwy 1 and 351, nr 351, W of Carberry, July 8, 1990, M.D. Schwartz, 2♀ (CNC). Carberry, June 14, 1953, A.R. Brooks, L.A. Kelton, 23, 5♀ (CNC). Carberry, June 15, 1953, A.R. Brooks, L.A. Kelton, Artemisia sp. (Asteraceae) , 123, 10♀ (CNC). Carberry, June 16, 1953, A.R. Brooks, L.A. Kelton, 13 (CNC). Carberry, June 18, 1953, A.R. Brooks, L.A. Kelton, 33, 18♀ (CNC). Carberry, June 19, 1953, A.R. Brooks, L.A. Kelton, 1♀ (CNC). Carberry, July 30, 1953, A.R. Brooks, L.A. Kelton, 1♀ (CNC). Carberry, July 31, 1953, A.R. Brooks, L.A. Kelton, 13, 2♀ (CNC). Carberry, August 9, 1953, A.R. Brooks, L.A. Kelton, 1♀ (CNC). Carberry, August 10, 1953, A.R. Brooks, L.A. Kelton, 13, 3♀ (CNC). Goodlands, June 12, 1931, R.H. Handford, 1♀ (CNC). Melita, July 25, 1958, A.R. and J.E. Brooks, 1♀ (CNC). Millwood, July 19, 1954, A.R. Brooks, Wallis, 23, 1♀ (CNC). Morris, August 15, 1953, A.R. Brooks, 13, 5♀ (CNC). Ninnette, June 11, 1958, R.B. Madge, 1♀ (CNC). Ninnette, June 20, 1958, R.B. Madge, 1♀ (CNC). Oak Lake, July 9, 1953, A.R. Brooks, L.A. Kelton, 3♀ (CNC). Oak Lake, July 11, 1953, A.R. Brooks, L.A. Kelton, 53, 12♀ (CNC). Pilot Mound, July 31, 1958, A.R. and J.E. Brooks, 1♀ (CNC). Riding Mountain National Park, July 20, 1972, L.A. Kelton, 73, 6♀ (CNC). 10 mi W of Roblin, July 15, 1954, A.R. Brooks, L.A. Kelton, 13, 2♀ (CNC). Roblin, July 15, 1954, A.R. Brooks, L.A. Kelton, 13 (CNC). Russell, July 21, 1973, A.R. Brooks, L.A. Kelton, 1♀ (CNC). Russell, July 17, 1954, A.R. Brooks, Wallis, 23, 8♀ (CNC). Russell, July 31, 1973, L.A. Kelton, Artemisia dentata (Asteraceae) , 6♀ (CNC). Saint Lazare, August 19, 1954, A.R. Brooks, L.A. Kelton, 23, 3♀ (CNC). 5 mi SW of Shilo, June 28, 1958, C.D.F. Miller, 5♀ (CNC). 8 km E of Roblin on Rt 5, August 21, 1993, M. D. Schwartz, Artemisia frigida (Asteraceae) , 203 (CNC). 8 km N of Spirit Sands, Spruce Wood Prov. Park (16 km S of Carberry), July 8, 1990, M. D. Schwartz, Artemisia frigida (Asteraceae) , 593 (CNC). Treesbank, July 7, 1928, R. H. Handford, 1♀ (CNC). Virden, July 8, 1953, A.R. Brooks, L.A. Kelton, 33, 6♀ (CNC). Virden, July 9, 1953, A.R. Brooks, L.A. Kelton, 13, 1♀ (CNC). Virden, July 10, 1953, A.R. Brooks, L.A. Kelton, 43, 9♀ (CNC). Virden, July 13, 1953, A.R. Brooks, L.A. Kelton, Artemisia frigida (Asteraceae) , 13♀ (CNC). 4.5 mi E of Rt 201 on Rt 59, July 7, 1990, M. D. Schwartz, Artemisia frigida (Asteraceae) , 343 (CNC). Northwest Territories: Arctic River Road, July 18, 1980, R. J. Cannings, 23, 4♀ (UBC). Inuvik, July 19, 1980, R. J. Cannings, 2♀ (UBC). Ontario: Sandbanks Provincial Park, August 13, 1991, M.D. Schwartz, Ambrosia trifida L. var. trifida (Asteraceae) , 13♀ (CNC). Quebec: Bradore Bay, August 2, 1930, W.J. Brown, 173, 22♀ (CNC). Bradore Bay, August 8, 1930, W.J. Brown, 53 (CNC). Choisy, nr Rigaud, August 25, 1956, Lindberg, 73, 16♀ (CNC). Great Whale River, July 22, 1949, J.R. Vockeroth, Achillea millefolium (Asteraceae) , 13 (CNC). Great Whale River, July 28, 1949, J.R. Vockeroth, 1♀ (CNC). Great Whale River, July 31, 1949, J.R. Vockeroth, 13 (CNC). Great Whale River, August 24, 1949, J.R. Vockeroth, Tanacetum huronense (Asteraceae) , 123, 31♀ (CNC). Laniel, July 20–21,1963, W. Gagne, 1♀ (CNC). Laniel, July 24–25, 1963, W. Gagne, 1♀ (CNC). Rupert House, August 3, 1949, D.P. Gray, 1♀ (CNC). Saskatchewan: Assiniboia, June 20, 1955, A.R. Brooks, 11♀ (CNC). Attons Lake, Cut Knife, July 11, 1940, A.R. Brooks, 2♀ (CNC). Beaver Creek, September 8, 1950, L.A. Konotopetz, 23, 2♀ (CNC). Big River, July 5, 1959, A.R. and J.E. Brooks, 23, 3♀ (CNC). Buffalo Pound, June 29, 1955, A.R. Brooks, 2♀ (CNC). Canora, July 11, 1954, A.R. Brooks, Wallis, 23, 7♀ (CNC). Chaplin, June 21, 1954, A.R. Brooks, Wallis, 1♀ (CNC). Christopher Lake, July 11, 1959, A.R. and J.E. Brooks, 13 (CNC). Christopher Lake, July 13, 1959, A.R. and J.E. Brooks, 3♀ (CNC). Christopher Lake, July 15, 1959, A.R. and J.E. Brooks, 13, 1♀ (CNC). Cypress Hills, September 19, 1951, L.A. Konotopetz, 3♀ (CNC). Duck Lake, 16.3 km S of MacDowell Rd., July 12, 1990, M.D. Schwartz, 1♀ (CNC). Elbow, August 15, 1951, A.R. Brooks, 83, 3♀ (CNC). Elbow, June 20, 1954, A.R. Brooks, 1♀ (CNC). Elbow, June 23, 1954, A.R. Brooks, Wallis, 13 (CNC). Elbow, June 24, 1954, A.R. Brooks, Wallis, 2♀ (CNC). Elbow, June 17, 1960, A.R. Brooks, 23, 2♀ (CNC). Esterhazy, July 26, 1954, A.R. Brooks, Wallis, 3♀ (CNC). Esterhazy, August 14, 1954, A.R. Brooks, Wallis, 33, 7♀ (CNC). Estevan, June 15, 1955, A.R. Brooks, 1♀ (CNC). Gascoigne, August 23, 1957, A.R. and J.E. Brooks, 1♀ (CNC). Glidden, July 3, 1952, L.A. Konotopetz, 23 (CNC). Good Spirit Lake, July 10, 1954, A.R. Brooks, Wallis, 11♀ (CNC). Good Spirit Lake, August 23, 1954, A.R. Brooks, Wallis, 1♀ (CNC). Great Sand Hills, July 4, 1952, A.R. Brooks, 1♀ (CNC). Hudson Bay, July 9, 1954, A.R. Brooks, Wallis, 13, 6♀ (CNC). Kandahar, July 29, 1954, A.R. Brooks, Wallis, 14♀ (CNC). Kenosee, May 30, 1958, A.R. Brooks, 1♀ (CNC). Kenosee, June 7, 1958, A.R. Brooks, 2♀ (CNC). Lebret, July 5, 1951, L.A. Konotopetz, 43, 1♀ (CNC). Lisieux, June 21, 1955, A.R. Brooks, 1♀ (CNC). Lumsden, August 7, 1954, A.R. Brooks, Wallis, 53, 2♀ (CNC). Lumsden, July 12, 1958, A.R. and J. Brooks, Wallis, 1♀ (CNC). 10.8 km E of Maidstone on Hwy 16, August 22, 1993, M. D. Schwartz, Artemisia cana (Asteraceae) , 23, 2♀ (CNC). McGee, July 3, 1952, L.A. Konotopetz, 1♀ (CNC). Minton, August 11, 1955, A.R. Brooks, 2♀ (CNC). Mossbank, June 23, 1955, A.R. Brooks, 5♀ (CNC). Pike Lake, September 1, 1949, A.R. Brooks, 43, 6♀ (CNC). Pike Lake, June 14, 1950, B.P. Beirne, 1♀ (CNC). Prince Albert, July 3, 1954, A.R. Brooks, Wallis, 3♀ (CNC). Prince Albert, July 4, 1954, A.R. Brooks, Wallis, 7♀ (CNC). Prince Albert, July 5, 1954, A.R. Brooks, Wallis, 13, 2♀ (CNC). Prince Albert, July 23, 1959, A. and J. Brooks, 4♀ (CNC). Rockglen, June 21, 1955, A.R. Brooks, 1♀ (CNC). Rockglen, August 2, 1955, A.R. Brooks, 1♀ (CNC). Rutland, July 23, 1940, A.R. Brooks, 6♀ (CNC). Rutland, August 2, 1940, A.R. Brooks, 4♀ (CNC). Saint Victor, June 25, 1955, A.R. Brooks, Wallis, 13, 2♀ (CNC). Saskatoon, June 13, 1950, A.R. Brooks, Artemisia sp. (Asteraceae) , 1♀ (CNC). Saskatoon, June 20, 1949, L.A. Konotopetz, Artemisia sp. (Asteraceae) 1♀ (CNC). Saskatoon, June 29, 1923, K. M. King, 13 (CNC). Saskatoon, July 17, 1924, K. M. King, 1♀ (CNC). Saskatoon, July 21, 1950, A.R. Brooks, Artemisia sp. (Asteraceae) , 9♀ (CNC). Saskatoon, July 22, 1950, A.R. Brooks, Artemisia frigida (Asteraceae) , 1♀ (CNC). Saskatoon, August 10, 1950, A.R. Brooks, Artemisia sp. (Asteraceae) , 1♀ (CNC). Saskatoon, August 11, 1949, A.R. Brooks, Artemisia sp. (Asteraceae) , 2♀ (CNC). Saskatoon, August 15, 1950, A.R. Brooks, Artemisia sp. (Asteraceae) , 13, 1♀ (CNC). Saskatoon, August 25, 1923, K. M. King, 1♀ (CNC). Saskatoon, August 27, 1923, K. M. King, 1♀ (CNC). Saskatoon, September 5, 1926, K. M. King, 1♀ (CNC). Saskatoon, September 6, 1923, K. M. King, 1♀ (CNC). Saskatoon, September 14, 1949, A.R. Brooks, L. A. Konotopetz, Artemisia sp. (Asteraceae) , 13, 51♀ (CNC). Saskatoon, September 19, 1923, K. M. King, 2♀ (CNC). Saskatoon, September 27, 1923, K. M. King, 13 (CNC). Saskatoon, September 30, 1923, K. M. King, 33 (CNC). 45.8 km N of Stoughton on Rt 47, July 9, 1990, M. D. Schwartz, Artemisia frigida (Asteraceae) , 303 (CNC). Swift Current, August 30, 1939, A.R. Brooks, 2♀ (CNC). Swift Current, September 1, 1939, A.R. Brooks, 13, 6♀ (CNC). Torch River, August 23, 1950, L.A. Konotopetz, 13 (CNC). 10.7 km S of Torch River Campground, on Rt 106, July 12, 1990, M.D. Schwartz, 13 (CNC). Torquay, August 14, 1955, A.R. Brooks, 13 (CNC). Tunstal, August 27, 1957, A.R. and J.E. Brooks, 13, 3♀ (CNC). Val Marie, June 12, 1955, A.R. Brooks, 2♀ (CNC). Val Marie, August 8, 1955, A.R. Brooks, 13, 3♀ (CNC). Val Marie, August 10, 1955, C. D. Miller, 13 (CNC). Willow Bunch, July 20, 1955, C. D. Mill­ er, 13, 2♀ (CNC). Willow Bunch, July 27, 1955, A.R. Brooks, 13, 3♀ (CNC). Willow Bunch, July 29, 1955, A.R. Brooks, 13 (CNC). Willows, June 19, 1955, A.R. Brooks, 13, 5♀ (CNC). Wilczek Lake, July 19, 1982, G.G.E. Scudder, 23 (UBC). Wood Mountain, June 17, 1955, A.R. Brooks, 13 (CNC). Yukon Territory: Aishihik River, Alaska Hwy, August 13, 1983, J. J. Robinson, Artemisia frigida (Asteraceae) , 23 (UBC). Burwash Landing, July 8, 1983, J. J. Robinson, 13 (UBC). Canyon, July 9, 1983, G.G.E. Scudder, 23 (UBC). Carcross, July 29, 1982, G.G.E. Scudder, Artemisia frigida (Asteraceae) (UBC). Carcross, August 9, 1983, L.A. Kelton, Artemisia frigida (Asteraceae) , 49♀ (CNC). Carmacks, July 17, 1983, L.A. Kelton, Artemisia sp. (Asteraceae) , 59♀ (CNC). Carmacks, July 28, 1982, G.G.E. Scudder, 23 (UBC). Dawson, July 23, 1983, L.A. Kelton, Artemisia frigida (Asteraceae) , 1♀ (CNC). Dawson, July 11, 1983, G.G.E. Scudder, 23 (UBC). Dawson, July 21, 1982, L.A. Kelton, Artemisia frigida (Asteraceae) , 113, 3♀ (CNC). Destruction Bay, July 26, 1982, L.A. Kelton, Salix sp. (Saliaceae) , 1♀ (CNC). Engineer Creek, Dempster Highway, July 27, 1983, L.A. Kelton, 3♀ (CNC). Frog Lake, July 13, 1981, S. G. Cannings, 33 (UBC). Koidern (CNC). Gravel Lake, 58 mi E of Dawson, 2050 ft, August 11, 1962, P.J. Skisko, 33, 7♀ (CNC). Haines Junction, July 28, 1982, L.A. Kelton, 13, 1♀ (CNC). Junction of Dempster and Klondike Highway, July 20, 1982, L.A. Kelton, 23 (CNC). Koidern, July 24, 1982, L.A. Kelton, 63, 2♀ (CNC). 10 km E of McQuesten, July 13, 1980, C. Guppy, Artemisia frigida (Asteraceae) , 33 (UBC). Moose Creek, July 19, 1982, G.G.E. Scudder, 33 (UBC). Moose Creek, July 18, 1982, L.A. Kelton, 313, 39♀ (CNC). Morley River, August 9, 1982, L.A. Kelton, 13, 3♀ (CNC). Old Crow, June 30, 1983, S. G. Cannings, 133 (UBC). Pelly Crossing, July 29, 1983, L.A. Kelton, Artemisia sp. (Asteraceae) , 11♀ (CNC). Pelly Crossing, July 2, 1985, E. Bijdemast, 33 (UBC). Porcupine River, Pine Bluffs, July 11, 1982, C. Guppy, Artemisia frigida (Asteraceae) , 113 (UBC). Rancheria, August 11, 1982, L.A. Kelton, Lathyrus sp. (Fabaceae) , 13 (CNC). Silver City, July 27, 1982, L.A. Kelton, 13, 1♀ (CNC). Snag Junction, July 23, 1982, L.A. Kelton, 23, 5♀ (CNC). Stewart Crossing, July 27, 1982, G.G.E. Scudder, 93 (UBC). Tagish, August 3, 1982, L.A. Kelton, Shepherdia canadensis (Elaeagnaceae) , 13, 3♀ (CNC). Takhini Hot Springs, 2400 ft, August 19, 1962, R.E. Leech, Artemisia sp. (Asteraceae) , 4♀ (CNC). Tatchum, July 19, 1982, G.G.E. Scudder, 43 (UBC). Tatchun, August 11, 1983, L.A. Kelton, Artemisia frigida (Asteraceae) , 23, 16♀ (CNC). Von Wilczek, July 18, 1982, L.A. Kelton, 93, 9♀ (CNC). Whitehorse, July 16, 1982, L.A. Kelton, Artemisia frigida (Asteraceae) , 53 (CNC). Whitehorse, July 30, 1982, L.A. Kelton, Artemisia frigida (Asteraceae) , 13, 5♀ (CNC). Whitehorse, August 6, 1949, L. C. Curtis, 13 (CNC). Whitehorse, August 19, 1959, R.B. Madge, 1♀ (CNC). MEXICO: Baja California Norte: 12 mi SE of Maneandero, April 19, 1965, Cavagnaro, Rose, Vesterby, 13 (CAS). 18.2 mi E of San Telmo, May 23, 1980, Brown and Faulkner, 13 (SDNM). 19 km E of El Rosario, March 28, 1980, J. Pinto, 13 (UCR). Tecate, 0.3 mi E of El Condor, April 15, 1982, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 13 (AMNH). Durango: Durango, July 4, 1964, L.A. Kelton, 53, 2♀ (CNC). USA: Alaska: Anchorage, August 5, 1954, A. G. Hunt, 13 (KU). Anchorage, Fish Creek Flats, July 27, 1948, R. I. Sailer, 23 (USNM). Chicken, July 22, 1982, L.A. Kelton, Betula glandulosa (Betulaceae) , 13, 3♀ (CNC). Copper Center, July 23, 1948, F. S. Blanton, Aster sp. (Asteraceae) , 23, 4♀ (USNM). Dyea, July 6, 1980, R. J. Cannings, 23, 1♀ (UBC). Lawing, July 22, 1951, W.J. Brown, on tundra, 13 (CNC). Skagway, July 5, 1980, R. J. Cannings, 1♀ (UBC). Naknek, August 9, 1952, W.R.M. Mason, 13, 1♀ (CNC). Skagway, September 6, 1936, S. M. Williams, 13 (AMNH). Arizona: Apache Co. : Alpine, September 1, 1940, L. L. Stitt, 23, 2♀ (USNM). Coconino Co.: Flagstaff, 7000 ft, May 8, 1967, D. C. and K. A. Rentz, 63 (UCB). Flagstaff, August 5, 1933, R. H. Beamer, 33 (KU). Grand Canyon Natl. Park, July 15, 1947, R. H. Beamer, 33 (KU). Grand Canyon, 7000 ft, June 22, 1925, A. A. Nichol, 83, 8♀ (USNM). Grand Canyon, June 29, 1930, R. L. Usinger, Artemisia sp. (Asteraceae) , 103 (UCB). Grand Canyon, South Rim, June 29, 1930 – June 16, 1949, R. L. Usinger, G. F. Knowlton, Artemisia sp. (Asteraceae) , 223 (USU,UCB). Parks, Kaibab National Forest, August 16, 1967, L.A. Kelton, 13 (CNC). Oak Creek Canyon, July 9, 1941 – July 9, 1942, Burt Hodgen, Beamer, 43 (KU). Rt 64 just SE of Grand Canyon Natl. Park in Kaibab National Forest, June 26, 1980, K. and R. Schmidt, Artemisia sp. (Asteraceae) , 103 (AMNH). Sunset Crater Road, 1 mi E of Rt 89A, 7300 ft, June 16, 1983, R. T. Schuh, M. D. Schwartz, Artemisia sp. (Asteraceae) , 723 (AMNH). Williams, June 24, 1925, A. A. Nichol, 33 (USNM). Graham Co.: Graham Mts., mp 120 on Rt 336, 4600 ft, April 25, 1982, M. D. Schwartz, Artemisia dracunculus (Asteraceae) , 313 (AMNH). Greenlee Co.: Apache National Forrest on Hwy 78, 5.4 mi SW Arizona – New Mexico state line, September 20, 1998, J. C. Schaffner, 933 (TAMU). Navajo Co.: Show Low, Sitgreaves National Forest, August 10, 1967, L.A. Kelton, 93, 9♀ (CNC). Pima Co. : Santa Catalina Mts. , May 31, 1957, Butler and Werner, 13 (UAZ). Mt. Lemon, April 29, 1948, R. H. Beamer, 13 (KU). Yavapai Co: Prescott, August 16, 1972, L.A. Kelton, 13 (CNC). California: Alameda Co.: East Oakland, February 22, 1936, P. Kinstry, 13 (KU). Niles Canyon, June 15, 1916 – May 26, 1928, E. P. Van Duzee, R. L. Usinger, 53 (CAS,UCB). Niles Canyon, May 26, 1928, R. L. Usinger, 43 (UCB). Redwood Canyon, July 1, 1916, W. M. Giffard, 13 (CAS). Contra Costa Co.: Mt. Diablo, July 21, 1935, R. H. Beamer, 23 (KU). San Pablo, July 26, 1969, Gagne, 13 (UCB). El Dorado Co.: Lake Tahoe, August 11, 1940, L. J. Lipovsky, 13 (KU). Road to Fallen Leaf Lake, 6300 ft, August 28, 1981, K. A. Phillips, 63 (OSU). Inyo Co. : 4 mi NW of Independence, June 2, 1976, J. Pinto, 13 (UCR). 7 mi N of Parchers Camp, June 30, 1961, J. K. Drew, 13 (CAS). E side of California Rt 141, 4000 ft, May 31, 1981, J. T. Polhemus, 33 (JTP). Independence, June 14, 1929, R. L. Usinger, 33 (CAS). Mammoth Lake, July 29, 1940, D. E. Hardy, 73 (KU). Lassen Co.: Doyle State Wildlife Preserve, June 24, 1974, Oman, 13 (OSU). Litchfield, June 22, 1937, B. P. Bliven, 13 (CAS). Sage Hen, July 27, 1954 – August 20, 1969, B. P. Bliven, W. Gagne, 43 (CAS,UCB). Los Angeles Co.: Malibu, February 27, 1966, 23 (LACM). 11.5 mi W of Malibu, April 15, 1965, F. Rindge, 13 (AMNH). 2 mi N of Escondido Beach, March 16, 1967, J. Powell, 53 (UCB). Claremont, Baker [no date], 23 (AMNH). Claremont, Baker [no date], 63 (HELSINKI). Latigo Canyon Road, 11.6 mi WNW of Malibu, April 15, 1965 – June 19, 1975, M. W. Hetz, F. J. & M. Rindge, 23 (UAZ, AMNH). Los Angeles, Coquillett, 53, 5♀ (USNM). Palos Verdes, Sea Side, March 24, 1981, J. T. Polhemus, 93 (JTP). San Fernando, February 23, 1941, Dale Lindsay, 13 (KU). San Vicente, May 14, 1938, C. E. Norland, 23 (LACM). West Fork San Gabriel River, May 5, 1996, F. W. Furry, 13 (LACM). Westwood Hills, May 11, 1942 – December 20, 1962, R. Beer, C. Ishida, 33 (LACM, KU). Whittier, March 24, 1916, L. L. Muchmore, 43 (LACM). Marin Co.: Muir Woods, June 6, 1950, W. J. Wall, 13 (UCD). Stinson Beach, August 15, 1938, R. H. Beamer, 13 (KU). Mariposa Co. : Yosemite National Park, August 4, 1940, R. H. Beamer, 13 (KU). Modoc Co.: 1 mi S of Lava Beds Natl. Mon. on Hill Rd, Modoc National Forest, July 17, 1985, G. M, Stonedahl, J. D. McIver, Artemisia tridentata (Asteraceae) , 43 (AMNH). 2.5 mi S of Rt 139 towards Lookout, 1440 m, July 6, 1979, R. T. and Joe Schuh, Artemisia sp. (Asteraceae) , 13 (AMNH). 24.7 mi NW of Canby, 1375 m, July 1, 1979, R. T. Schuh and B. M. Massie, 23 (AMNH). 4 mi E of Cedarville, 1440 m, July 2, 1979, R. T. Schuh and B. M. Massie, 13 (AMNH). Mono Co.: Coleville, June 10, 1966, W. Gagne, 13 (UCB). Crooked Creek Naval Res. Station, White Mts. , 10,000 ft, August 4, 1966, L. and C. W. O’Brien, 113 (UCB). Mammoth Camp, W. D. Pierce, 23 (LACM). McKay Creek, Sonora Pass, August 18, 1960, E Jessen, 13 (UCB). Mono Lake at Hwy 395, 2188 m, July 11, 1980, R. T. Schuh, G. M. Stonedahl, Artemisia sp. (Asteraceae) , 23 (AMNH). Mono Lake, January 1, 1939, W. D. Pierce, 23 (LACM). Nevada Co.: Hobart Mills, Prosser Creek, October 10, 1952, J. C. Hall, 13 (UCD). Sagehen Creek Station near Hobart Mills, August 20, 1969, W. Gagne, Artemisia sp. (Asteraceae) , 43 (UCB). Orange Co.: Black Star (Canyon?), March 10, 1935, 13 (LACM). Green River Camp, Lower Santa Ana Canyon , May 9, 1933, E. P. Van Duzee, 43 (CAS). Newport Beach, April 24, 1971, Oman, 23 (OSU). Placer Co.: Truckee, Tahoe National Forest, August 29, 1967, L.A. Kelton, Artemisia sp. (Asteraceae) , 223, 8♀ (CNC). Plumas Co.: near Chester, July 5, 1932, R. L. Usinger, 13 (UCB). Riverside [?] Co.: Peralta, May 7, 1948, R. A. Flock, Artemisia vulgaris (Asteraceae) , 13 (UCR). Riverside Co.: 5.5 SE of Sage, March 14, 1979, J. Pinto, 13 (UCR). Hwy 74, Cleveland National Forest, El Cariso Station, 2500 ft, May 12, 1978, J. Pinto, Artemisia sp. (Asteraceae) , 33 (UCR). Lower Tin Mine Canyon, July 14, 1963, M. Irwin, 33 (UCR). Menifee Valley, hills on W end, March 10, 1979, J. Pinto, 33 (UCR). Palm Springs, December 22, 1941, R. H. Beamer, 23 (KU). Quail Valley, May 20, 1974 – May 2, 1977, J. Pinto, 163 (UCR). Riverside, March 26, 1989, R. Wharton, 13 (TAMU). Riverside, March 29, 1971 – March 26, 1989, E. J. Paddock, 13 (CAFA). Saboba Springs, July 4, 1917, E. P. Van Duzee, 53 (CAS). University of California at Riverside, June 14, 1982, T. J. Henry, Artemisia californica (Asteraceae) , 33, 6♀ (USNM). Sacramento Co.: Sacramento, June 3, 1920 – August 10, 1935, H. H. Keifer, 23 (CAFA). San Bernardino Co.: Cajon Pass Summit on I­15, July 2, 1949, Flock, 13 (UCR). Colton, May 26, 1917 – May 28, 1917, E. P. Van Duzee, 43 (CAS). San Diego Co.: 8.2 mi E of Dulzura on Rt 94 at mp 36.50, April 15, 1982, M. D. Schwartz, 13 (AMNH). Alpine, July 9, 1929, R. H. Beamer, 43 (KU). Dictionary Hill, December 4, 1979, Faulkner, Brown, 13 (SDNM). Mission Dam, January 2, 1928 – March 16, 1929, C. C. Searl, 233 (SDNM). Morena Dam, May 26, 1929, 13 (SDNM). Ramona, April 18, 1938, E. P. Van Duzee, 13 (CAS). San Diego, April 11, 1914 – July 24, 1941, E. P. Van Duzee, R. H, Beamer, 163 (CAS, KU). San Diego, April 24, 1920, E. P. Van Duzee, 43 (CAS). Sentanac Canyon, May 4, 1968, J. DeVol, 13 (UCR). San Joaquin Co.: Stockton, August 21, 1919, E. P. Van Duzee, 13 (CAS). San Luis Obispo Co.: Baywood Park, May 17, 1980, J. Pinto, Artemisia sp. (Asteraceae) , 103 (UCR). California Polytechnic State University Campus, Poly Canyon, 125 m, May 8, 1985, R. T. Schuh and B. M. Massie, Artemisia californica (Asteraceae) , 163 (AMNH). Dune Lakes, May 20, 1972, J. D. Pinto, 13 (UCR). San Luis Obispo, April 24, 1919, E. P. Van Duzee, 63 (CAS). Santa Barbara Co. : 23 mi E of Santa Maria , June 20, 1952, R. E. and L. D. Beamer, 13 (KU). 6 mi SE of Lompoc, March 14, 1967, P. A. Opler, 33 (UCB). Jalama Beach, July 9, 1965, M. R. Gardiner, 23 (UCD). Santa Barbara , June 16, 1951, K. W. Tucker, 33 (UCD). Santa Cruz Island, Canyon Del Medio, March 2, 1971, R. O. Schuster, 13 (UCD). Santa Cruz Island, Puerto Canyon, March 21, 1977, A. J. Mayor, Artemisia californica (Asteraceae) , 153 (UCR). Upper Oso Campground off Rt 154, 310 m, May 7, 1985, R. T. Schuh, Artemisia californica (Asteraceae) , 93 (AMNH). Santa Clara Co. : Pacheco Pass on Rt 152, June 29, 1965, G. F. Knowlton, 13 (USU). Shasta Co. : Hat Creek, July 2, 1955, E. E. Lindquist, 13 (UCB). Sierra Co. : Sattley, July 7, 1970, R. M. Bohart, 13 (UCD). Siskiyou Co.: 1– 2 mi N of Tule Lake Refuge Hdq, 1255 m, June 26, 1979, R. T. and Joe Schuh, Artemisia tridentata (Asteraceae) , 33 (AMNH). 12.3 mi N of St. Hwy. 89 on Powder Hill Road, July 19, 1985, G. M, Stonedahl, J. D. McIver, Chrysothamnus viscidiflorus (Asteraceae) , 343 (AMNH). 15 mi SE of Mt. Shasta , 3500 ft, July 10, 1972, J. D. Lattin, 23 (OSU). 2 mi N of Tule Lake Refuge Hdq, 4000 ft, June 26, 1979, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 63 (AMNH). 2.5 mi N of Medicine Lake on Medicine Lake Rd, July 18, 1985, G. M. Stonedahl and J. D. McIver, Chrysothamnus sp. (Asteraceae) , 573 (AMNH). Bray, June 30, 1935, R. H. Beamer, 33 (KU). Yreka, June 15, 1959, L.A. Kellton, R. Madge, 13, 1♀ (CNC). Sonoma Co. : Guernwood Pk., May 30, 1928, R. L. Usinger, 13 (UCB). No specific locality, 13 (CAS). Trinity Co.: Coffee Creek, July 14, 1955, D. L. Dahisten, 43 (UCD). Tulare Co.: 2.6 mi W of county line near Chimney Peak RS, 2000 m, July 1, 1980, R. T. Schuh, 13 (AMNH). 21 mi W of Rt 395 toward Kennedy Meadows, 1875 m, July 1, 1980, R. T. Schuh, Artemisia tridentata (Asteraceae) , 53 (AMNH). Mineral King, July 29, 1935, Graham Heid, 13 (CAS). Ventura Co.: Casitas Reserve, N end, March 15, 1967, J. Powell, 83 (UCB). Oak View, December 30, 1964, P. M. Jump, 23 (LACM). Ventura, January 20, 1965, P. M. Jump, 53 (LACM). Ventura, January 23, 1919 – January 20, 1965, P. M. Jump, E. O. Essig, 153 (LACM, UCB). Ventura, January 23, 1919, E. O. Essig, Artemisia californica (Asteraceae) , 33 (UCB). Colorado: Archuleta Co.: Pagosa Springs, 7200 ft, August 24, 1972, L.A. Kelton, 13, 4♀ (CNC). Boulder Co.: 6 mi E of Nederland, August 8, 1973, J. C. Schaffner, 13 (TAMU). 6 mi W of Boulder, August 12, 1973, J. C. Schaffner, 53 (TAMU). 5 mi S of Nederland, 8000 ft, July 2, 1961, J.G. Chillcott, 1♀ (CNC). Bourbon Co.: No specific locality, 800 ft, January 1, 1915, R. H. Beamer, 13 (KU). Clear Creek Co.: Mount Evans, 13,000 ft, July 13, 1961, C.H. Mann, 1♀ (CNC). Mt. Goliath, 12,000 ft, August 7, 1987, D. A. and J. T. Polhemus, 23 (JTP). Mt. Goliath, 12,000 ft, June 21, 1988, J. T. Polhemus, 23 (JTP). West Chicago Creek, Arapaho National Forest, 9800 ft, July 28, 29, 1968, L.A. Kelton, 7 ♀ (CNC). Conejos Co.: Elk Creek Recreation Area, August 22, 1969, J. C. Schaffner, 13 (TAMU). Denver Co.: Denver, July 12, 1900 – July 12, 1903, Van Duzee, 33 (CAS). Dolores Co.: 29 mi SW Norwood, July 7, 1980, J. T. and D. A. Polhemus, 23 (JTP). Douglas Co.: 3 mi E of Larkspur, July 3, 1979, D. A. Polhemus, 33 (JTP). Perry Park, July 13, 1977, D. A. and J. T. Polhemus, 23 (JTP). Roxborough Road near Chatfield Park, May 28, 1978, J. T. Polhemus, 73 (JTP). Waterton, Head of Hiline, June 9, 1978 – August 7, 1981, J. T. Polhemus, 263 (JTP). Waterton, June 9, 1978 – August 7, 1981, D. A. Polhemus, Artemisia sp. (Asteraceae) , 383 (JTP). Eagle Co.: 9.2 mi S of Minturn, White River Natl. Forrest , 9080 ft, July 19, 1990, Zolnerowich, 43 (TAMU). Near Bond, Water Wheel Ranch, June 24, 1978, J. T. Polhemus, Artemisia tridentata (Asteraceae) , 63 (JTP). El Paso Co.: Calhan, August 19, 1936, W. D. Field, 13 (KU). Garden of Gods, June 16, 1968, K. E. Slansky, 13 (UCD). Elbert Co.: 3 mi E of Kiowa, 6500 ft, July 16, 1983, R. T. Schuh, D. A. and J. T. Polhemus, 393 (AMNH). 5 mi E of Agate, August 9, 1987, J. T. and D. A. Polhemus, Artemisia filifolia (Asteraceae) , 33 (JTP). Garfield Co.: W. Evacuation Creek, July 9, 1981, M. D. Schwartz, 13 (AMNH). Gilpin Co.: Aspen, White River National Forest, August 24, 1968, L.A. Kelton, Artemisia sp. (Asteraceae) , 83, 6♀ (CNC). Pinecliffe, August 12, 1973, J. C. Schaffner, 13 (TAMU). Pinecliffe, July 19, 1949 – August 12, 1973, J. R. White, J. C. Schaffner , 113 (KU, TAMU). Rollinsville, Roosevelt National Forest, September 2, 1972, L.A. Kelton, Pinus flexilis (Pinaceae) 13, 2♀ (CNC). Gunnison Co.: 5 mi NE of Almont, along Taylor River, 8400 ft, August 31, 1980, D. A. and J. T. Polhemus, Artemisia filifolia (Asteraceae) , 143 (JTP). Taylor Park, July 25, 1982, J. T. Polhemus, 13 (JTP). Jackson Co.: Cowdrey, 8000 ft, August 15, 1968, L.A. Kelton, 1♀ (CNC). 4 mi N of Gould, August 5, 1975, J. C. Schaffner, 443 (TAMU). Gould, 9200 ft, August 13, 1968 – August 13, 1969, Oman, Schaffner, 23 (OSU, TAMU). Muddy Pass, Routt National Forest, Artemisia sp. (Asteraceae) (CNC). Gould, 9200 ft, August 12–13, 1968, L.A. Kelton, Artemisia sp. (Asteraceae) , 183, 24♀ (CNC). Grand Lake Entrance, Rocky Mountain National Park, August 18, 1968, L.A. Kelton, Artemisia sp. (Asteraceae) , 63, 6♀ (CNC). Muddy Pass, Routt National Forest, 8600 ft, August 19, 1968, L.A. Kelton, Artemisia sp. (Asteraceae) , 23, 4♀ (CNC). Muddy Pass, Routt National Forest, 8600 ft, August 21, 1968, L.A. Kelton, Artemisia sp. (Asteraceae) , 23, 3♀ (CNC). Jefferson Co.: 10 mi NW of Golden, Guy Gulch, 7300 ft, August 10, 1987, J. T. and D. A. Polhemus, 13 (JTP). Red Rocks Park near Morrison, July 15, 1983 – July 26, 1988, D. A. Polhemus, R. T. Schuh, 403 (AMNH, JTP). Red Rocks Park, July 26, 1988, D. A. Polhemus, Artemisia ludoviciana (Asteraceae) , 123 (JTP). Upper Beaver Br. Gulch, August 12, 1981, D. A. Polhemus, 23 (JTP). Lake Co.: 10 mi N of Leadville, Tennessee Pass, 10,400 ft, July 24, 1988, R. Wharton, 23 (TAMU). La Plata Co.: La Plata, San Juan National Forest, 8500 ft, July 19–21, 1972, L.A. Kelton, 1♀ (CNC). Larimer Co.: 46 mi W of Fort Collins, Bennett Creek Picnic Grounds., Pingree Park Road, 7400 ft, July 14, 1986, R. T. Schuh, J. T. Polhemus, 543 (AMNH). 46 mi W of Fort Collins, Bennett Creek Picnic Grounds., Pingree Park Road, 7700 ft, July 14, 1986, R. T. Schuh, Artemisia frigida (Asteraceae) , 303 (AMNH). 46 mi W of Fort Collins, Pingree Park Rd, 7700 ft, August 19, 1924 – July 14, 1986, R. T. Schuh, J. T. Polhemus, Beamer, Lawson, 343 (AMNH, KU). Estes Park, August 24, 1925, H. H. Knight, 53, 5♀ (USNM). Estes Park, July 10, 1964, H. H. Knight, 103, 10♀ (USNM). Estes Park, July 18, 1916 – August 27, 1920, H. C. Severin, 23 (KU, AMNH). Fort Collins, July 13, 1900, E. P. Van Duzee, 13 (CAS). Rocky Mountain National Park, Grand Lake Entrance, August 5, 1947, R. H. Beamer, 13 (KU). Rustic (CNC). Pingree Park, September 3, 1972, L.A. Kelton, 7♀ (CNC). Poudre Canyon, Rustic, September 6, 1972, L.A. Kelton, 23, 3♀ (CNC). Las Animas Co.: 1 mi E of Stonewall, Fire Dept., 7400 ft, August 18, 1986, R. T. Schuh, 1043 (AMNH). Logan Co.: 15 mi N of Sterling, June 20, 1979, D. A. Polhemus, 83 (JTP). Mesa Co. : 20 mi E of Gateway, Upsweep Canyon, 6800 ft, August 15, 1987, J. T. and D. A. Polhemus, 33 (JTP). Colorado National Monument, Liberty Cap Trailhead, 6000 ft, June 10, 1982, M. D. Schwartz, 13 (AMNH). Moffat Co.: Gates of Lodore, June 28, 1979, D. A. Polhemus, 23 (JTP). Montezuma Co. : Mesa Verde Natl. Park , July 13, 1930 –July 30, 0130, R. L. Usinger, 23 (UCB). Mesa Verde National Park, 7000 ft, July 17, 18, 1968, L.A. Kelton, 4♀ (CNC). Ouray Co.: Ouray, August 26, 1972, L.A. Kelton, 1♀ (CNC). Routt Co.: Steamboat Springs, 6900 ft, July 16, 1964, H. H. Knight, 53, 5♀ (USNM). Steamboat Springs, July 16, 1964, H. H. Knight, Artemisia sp. (Asteraceae) , 103, 10♀ (USNM). Steamboat Springs, 6700 ft, July 15, 1964, H. H. Knight, det. as Psallus artemisicola Knight by Knight, 13 (CNC). Saguache Co.: 1 mi E of North Pass, August 19, 1969, J. C. Schaffner, 133 (TAMU). Summit Co.: Dillon, August 29, 1972, L.A. Kelton, Artemisia sp. (Asteraceae) , 43, 6♀ (CNC). Dillon Reservoir near Dillon Dam, August 15, 1981, D. A. Polhemus, Artemisia sp. (Asteraceae) , 13 (JTP). Teller Co.: Big Spring Ranch, July 30, 1966, T. C. Emmel, 13 (LACM). Woodland Park, 8000 ft, June 21, 1966, J. A. Slater, 23 (AMNH). Weld Co.: 7 mi NE of Nunn, August 15, 1968, Oman, 193 (OSU). Connecticut: New Haven, May 30, 1911, B. H. Walden, 1♀ (USNM). Idaho: Ada Co. : near Kuna, June 21, 1982, T. J. Henry, Artemisia sp. (Asteraceae) , 43, 3♀ (USNM). Bannock Co.: McCammon, June 29, 1966, W. Gagne, J. Haddock, 23 (UCB). Benewah Co.: vicinity of Emida, St. Joe National Forest, T42N R53W, 3000 ft, August 5, 1986, Schwartz and Stonedahl, 13 (AMNH). Blaine Co.: 20 mi N of Shoshone, Big Wood River, July 1, 1966, W. Gagne, J. Haddock, 13 (UCB). 6 mi NW of Carey, Little Wood River, June 30, 1966, W. Gagne, J. Haddock, 23 (UCB). Caribou Co.: Blackfoot Reservoir, June 28, 1949, G. Ruhr, 13 (AMNH). Cassia Co. : Black Pine Mountains, September 29, 1969, G. F. Knowlton, 13 (USU). Burley, July 11, 1932, D. E. Fox, 4♀ (USNM). Latah Co.: Moscow, June 14, 1936, T. A. Brindley, 13 (USNM). Madison Co.: Rexburg, June 22, 1935, C. F. Smith, 23 (USU). Nez Perce Co.: Lewiston, 550 ft, May 30, 1932, T. A. Brindley, 13 (USNM). Lewiston, May 30, 1932, T. A. Brindley, 23 (AMNH). Lewiston, 550 ft, May 30, 1932, T.A. Brindley, det. as Chlamydatus bakeri Bergroth by H.H. Knight, paratype of Psallus artemisicola Knight , 2♀ (CNC). Oneida Co. : 6 mi NW of Holbrook, June 13, 1969 – June 30, 1969, G. F. Knowlton, 133 (USU). Black Pine Canyon, 7000 ft, June 4, 1974, G. E. Bohart, 13 (USU). Ireland Canyon, July 11, 1973, G. F. Knowlton, 13 (USU). Juniper, September 29, 1969, G. F. Knowlton, 13 (USU). Pocatello Valley, July 11, 1973, G. F. Knowlton, 13 (USU). Salyer Cow Camp, July 23, 1971, W. J. Hanson, 13 (USU). Power Co.: American Falls, August 5, 1972, L.A. Kelton, Juniperus sp. (Cupressaceae) , 13, 2♀ (CNC). Twin Falls Co.: Hollister (plot 6 outside), June 13, 1931, D. E. Fox, 103, 10♀ (USNM). Unknown Co.: Giveout, July 7, 1920, 13 (AMNH). Tuttle, June 8, 1930, Harold Waters, 33, 3♀ (USNM). Illinois: Ogle Co.: Grand Detour, Castle Rock, July 12, 1934, Dozier and Mohr, 13 (INHS). Oregon, July 4, 1932, Dozier and Mohr, Artemisia canadensis (Asteraceae) , 23, 2♀ (USNM). Oregon, July 4, 1932, Dozier and Mohr, Artemisia sp. (Asteraceae) , 323 (INHS). Winnebago Co.: Rockton, July 5, 1932, Dozier and Mohr, 23 (INHS). Kansas: Chautauqua Co.: Castle Rock, July 8, 1937, C. L. Johnston, 13 (KU). Cedar Vale, June 23, 1943, R. H. Beamer, 33 (KU). Harvey Co.: Newton, August 3, 1945, R. H. Beamer, 43 (KU). Kearny Co.: Kearny, 2998 ft, July 19, 1921, 13 (KU). McPherson Co.: No specific locality, June 28, 1923, L. D. Anderson, 13 (KU). Michigan: Marquette Co.: Lakewood, August 18, 1960, L.A. Kelton, Whitney, Ambrosia sp. (Asteraceae) , 243, 20♀ (CNC). Minnesota: Cook Co.: Grand Marais, August 13, 1922, H. H. Knight, 153, 15♀ (USNM). Grand Marais, August 13, 1922, H.H. Knight, det. as Plagiognathus albipennis Fallen by H. H. Knight, 23 (CNC). Chisago Co.: North Branch, August 5, 1922, H.H. Knight, paratype of Psallus artemisicola Knight , 23, 2♀ (CNC). North Branch, August 5, 1922, H.H. Knight, det. as Chlamydatus bakeri Bergroth by H.H. Knight, paratype of Psallus artemisicola Knight , 33, 3♀ (CNC). Clay Co.: BB Ranch 4–5, T141N, R45W, sec. 8, August 4, 1995 – September 14, 1995, Rider, Fauske, Locken, 53, 12♀ (DAR). Bluestem Site, T139N, R46W, sec. 15, July 13, 1995 – August 22, 1995, Rider, Fauske, Locken, 6♀ (DAR). Clay County Land 2–3, T141N, R45W, August 15, 1995, D. Rider and C. Locken, 23 (DAR). Hennepin Co.: Fort Snelling, August 2, 1923, A. T. Hertig, 2♀ (USNM). Polk Co.: Agassiz Dunes, T147N, R44W, sec. 32, August 1, 1995, D. Marty and K. Urlacher, 13, 5♀ (DAR). Montana: Carbon Co.: Bear Creek between Red Lodge and Belfry, 5000 ft, August 12, 1986, Schuh, Schwartz, Stonedahl, Sarcobatus vermiculatus (Chenopodiaceae) , 63 (AMNH). Smith Mine, August 12, 1986, Schuh, Schwartz, Stonedahl, 33 (AMNH). Deer Lodge Co.: Silver Lake, 14 mi W of Anaconda on Rt 10A, August 9, 1986, Schuh, Schwartz, Stonedahl, 223 (AMNH). Gallatin Co.: Targhee Pass on Rt 191, 7072 ft, August 10, 1986, Schuh, Schwartz, Stonedahl, 63 (AMNH). 29 mi S of Bozeman Hot Springs on Rt 191, 5800 ft, August 10, 1986, Schuh, Schwartz, Stonedahl, 1043 (AMNH). Glacier Co.: 10 mi W of Babb, Glacier National Park, August 8, 1969, P. Oman, 33 (OSU). Nebraska: Brown Co.: Long Pine, July 15, 1986, T. J. Henry and A. G. Wheeler, Jr., Artemisia sp. (Asteraceae) , 103, 10♀ (USNM). Cherry Co.: Valentine, June 8, 1950 – June 9, 1950, Slater, Hicks, Laffoon, 73 (AMNH). Dodge Co.: Fremont, August 28, 1925, H. H. Knight, 13 (USNM). Nevada: Carson City Co.: Carson City, June 25, 1929 – May 30, 1947, R. L. Usinger, W. F. Chamberlain, 73 (CAS, TAMU). Carson City, June 25, 1929, E. P. Van Duzee, 53 (CAS). Elko Co.: 23.7 mi S of Hwy 229 on Hwy 93, 6400 ft, July 19, 1980, Stonedahl, Chrysothamnus nauseosus (Asteraceae) , 13 (AMNH). 30 mi SE of I­80 on Hwy 229, July 19, 1980, Stonedahl, Artemisia sp. (Asteraceae) , 23 (AMNH). 30 mi W of Elko on Rt 80, June 18, 1982, T. J. Henry, Artemisia sp. (Asteraceae) , 33, 8♀ (USNM). Carlin, June 25, 1935, R. H. Beamer, 13 (KU). Ruby Mts., Lomoille Canyon, June 16, 1983, R. T. Schuh and M. D. Schwartz, Artemisia dracunculus (Asteraceae) , 113 (AMNH). Utah state line on Utah Rt 30, June 25, 1983, R. T. Schuh, M. D. Schwartz, Artemisia spinescens (Asteraceae) , 23 (AMNH). Wells, July 12, 1965, H. H. Knight, 33, 3♀ (USNM). Esmeralda Co. : 3 mi W of Lida, June 7, 1966, W. Gagne, 23 (UCB). Eureka Co.: 12 mi N of Rt 50 on Rt 278, 5800 ft, June 27, 1983, R. T. Schuh, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 63 (AMNH). 28 mi W of Eureka on Rt 50, 6000 ft, June 17, 1983, R. T. Schuh and M. D. Schwartz, Artemisia tridentata (Asteraceae) , 13 (AMNH). 8 mi W Eureka, July 11, 1993, G. M. Chamberlain, 13 (TAMU). Lander Co.: 9.5 mi S Rt 2, Smith Creek Valley, T15N, R39E, June 30, 1983, R. T. Schuh, M. D. Schwartz, 53 (AMNH). Austin, August 12, 1940, L. J. Lipovsky, 13 (KU). Nye Co.: 11 mi N of Belmont, 7200 ft, June 29, 1983, R. T. Schuh and M. D. Schwartz, Artemisia nova (Asteraceae) , 183 (AMNH). 3.5 mi SE of Manhattan, Toiyabe National Forest, 2188 m, July 13, 1980, R. T. Schuh, G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 173 (AMNH). Mercury, 18 M, June 11, 1965, H. H. Knight and J. Merino, Artemisia tridentata (Asteraceae) , 13 (USNM). Mercury, 401M, June 18, 1965, H. H. Knight and J. Merino, 203, 20♀ (USNM). Mercury, 5 M, June 12, 1965, H.H. Knight, J. Merino, paratype of Europiella nigricornis Knight , 1♀ (CNC). Mercury, 18 M, June 11, 1965, H.H. Knight, J. Merino, Artemisia tridentata (Asteraceae) , paratype of Europiella nigricornis Knight , 23 (CNC). Mercury, 19 M, June 23, 1965, H.H. Knight, J. Merino, Artemisia tridentata (Asteraceae) , 13 (CNC). Ormsby Co.: No specific locality, July 1, 1900, Baker, 13 (CAS). Reno Co.: Reno, Sky Ranch, June 16, 1952, E. I. Schlinger, 33 (UCD). Washoe Co.: 13 mi SE of Reno, 5650 ft, Musgrave, 13 (OSU). 7 mi W of Vya on Rt 8A, 1813 m, July 2, 1979, R. T. Schuh and B. M. Massie, Artemisia tridentata (Asteraceae) , 223 (AMNH). Mt. Rose, July 9, 1970, Villegas, 13 (UCD). Patrick, June 16, 1964, P. Richerson, 13 (UCD). Reno, June 27, 1927, E. P. Van Duzee, 83 (CAS). Verdi, September 7, 1967, W. Gagne, 53 (UCB). White Pine Co. : 6 mi SW of Ely, June 23, 1966, W. Gagne and J. Haddock, 103 (UCB). Ely, July 3, 1960 – June 23, 1966, Tucker, Gagne, Haddock, 103 (UCD, UCB). Lehman Caves Natl. Mon., June 24, 1966, W. Gagne, J. Haddock, Artemisia sp. (Asteraceae) , 13 (UCB). New Mexico: Cantron Co.: 18 mi W Datil, August 15, 1982, J. C. Schaffner, 13 (TAMU). Catron Co.: 15.8 mi N of Silver City, 7160 ft, August 20, 1998, J. C. Schaffner, 13 (TAMU). 4 mi W of Mogollon, September 21, 1998, J. C. Schaffner, 223 (TAMU). Cibola Co.: 18 mi S El Malpais Natl. Conserv. Area, August 8, 1997, J. C. Schaffner, 23 (TAMU). Colfax Co. : 5 mi E of Eagles Nest, August 8, 1969, J. C. Schaffner, 73 (TAMU). Grant Co. : 3.4 mi W of Mule Creek, September 20, 1998, J. C. Schaffner, 23 (TAMU). Iron Creek CG , June 9, 1977 – June 4, 1980, Knowlton, Hanson, 73 (USU). Pinos Altos, August 19, 1972, L.A. Kelton, 13, 4♀ (CNC). Roberts Lake, Gila National Forest, August 19, 1972, L.A. Kelton, 83, 7♀ (CNC). Hidalgo Co.: Skeleton Canyon, August 16, 1968, G. C. Batista, 13 (UCR). Lea Co. : Site 12, September 24, 1979, R. R. Murray, J. C. Schaffner, 13 (TAMU). Lincoln Co.: Ruidoso, June 26, 1940, Hardy, Lipovsky, Beamer, Kuitart, Kenegan, 303 (KU). Sierra Blanca Mt. , 8600 ft, August 18, 1970, J. R. and M. H. Sweet, 53 (TAMU). Otero Co.: Mayhill, July 23, 1967, L.A. Kelton, 1♀ (CNC). 2 mi E of Mayhill, September 22, 1998, J. C. Schaffner, 103 (TAMU). Cloudcroft, June 17, 1976, P. Oman, 43 (OSU). Cloudcroft, June 27, 1940, R. H. Beamer, D. E. Hardy, Kenaga, Kuitert, 303 (KU). Mountain Park, June 27, 1940, L. J. Lipovsky, 33 (KU). Near Maryhill, Sacramento Mountains, September 1, 1998, M. D. Schwartz, 13, 1♀ (CNC). Rio Arriba Co. : 14 mi NW Chama, August 21, 1969, J. C. Schaffner, 93 (TAMU). Sandoval Co.: East Fork Jemez River, La Concha Campground, 8300 ft, August 14, 1972, R. E. Orth, 23 (UCR). Jemez Springs, August 23, 1972, L.A. Kelton, 73, 5♀ (CNC). Socorro Co. : No specific locality, August 18, 1927, Beamer, 13 (KU). Taos Co.: Taos, June 18, 1937, E. D. Ball, 13 (UAZ). Torrance Co.: Estancia, June 24, 1940, D. E. Hardy, 43 (KU). New York: Rockland Co.: South Nyack, Ross Avenue, June 11, 1988, M. D. Schwartz, 13 (AMNH). Suffolk Co.: Yaphank, May 29, 1982, A. G. Wheeler and T. J. Henry, Artemisia campestris (Asteraceae) , 83, 20♀ (PDA, USNM). Tompkins Co.: Ithaca, Cornell Plantations, August 15, 1987, A. G. Wheeler, Artemisia abrotanum (Asteraceae) , 63, 18♀ (PDA). North Dakota: Ada Co. : Leonard, July 26, 1937, R. H. Beamer, 23 (KU). McKenzie Co.: North Unit, Roosevelt National Memorial Park, June 30, 1973, L.A. Kelton, 23, 1♀ (CNC). Ransom Co.: 4 mi NW of McLeod, T134N, R53W, sec. 6, June 5, 1992, D. A. Rider, 1♀ (DAR). Richland Co.: 2.5 mi SE of McLeod, Cheyenne National Grassland, May 26, 1992, D. Rider, 13 (DAR). Trail Co.: Unknown locality, August 19, 1923, A.A. Nichol, paratype of Psallus artemisicola Knight , 13 (CNC). Unknown Co.: Beach, August 23, 1921, R. L. Webster, 1♀ (USNM). Oklahoma: Payne Co.: Stillwater, October 3, 1934, F. A. Fenton, 13 (USNM). Oregon: Baker Co.: 12 mi SW of Baker, June 20, 1968, Oman, 23 (OSU). Crook Co.: 23 mi E of Prineville, Mark’s Creek, July 29, 1976, J. D. Lattin, Artemisia arbustula (Asteraceae) , 23 (OSU). 3–15 mi SE of Prineville, July 2, 1968 – June 22, 1979, Oman, R. T. Schuh, M. D. Schwartz, 73 (OSU, AMNH). Just E of Prineville, June 22, 1979, R. T. Schuh, Artemisia tridentata (Asteraceae) , 83 (AMNH). Mitchell, July 22, 1979, G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 73 (AMNH). Deschutes Co.: 1 mi S of Millican, 4180 ft, June 21, 1979, R. T. Schuh, 13 (AMNH). 1 mi S of Millican, 4380 ft, June 7, 1977 – June 21, 1979, J. D. Lattin, R. T. Schuh, Artemisia tridentata (Asteraceae) , 63 (OSU, AMNH). 11 mi SE of Brothers, July 9, 1968, Lattin, Oman, 33 (OSU). 4 mi NE of Sisters, October 1, 1979, Lattin, 23 (OSU). Harney Co.: 3 mi S. Hdq of Squaw Butter Exp Sta., July 6, 1977, J. D. Lattin, 13 (OSU). Lily Lake, 13 mi E French Glen, July 10, 1968, Oman, 13 (OSU). T34S, R35E, sec. 10, off dunes, June 22, 1979, Lightfoot and Cobb, 13 (OSU). Jackson Co.: 1 mi below summit of Mount Ashland, 6500 ft, September 24, 1968, J. D. Lattin, 23 (OSU). 9 mi SE Butte Falls, July 19, 1973, Ritcher and Carlson, 43 (OSU). Ashland, August 2, 1918, E. P. Van Duzee, 53 (CAS). Jefferson Co.: 2.5 mi N of Warm Springs, 1700 ft, June 6, 1972, J. D. Lattin, 43 (OSU). Allen Spring, Metolius River, July 27, 1960, K. M. Fender, 13 (OSU). Klamath Co.: 1 mi E of Dairy, October 10, 1957, J. Lattin, 13 (OSU). 10 mi SE of Chiloquin, July 4, 1982, G. M, Stonedahl, T. J. Henry, Artemisia tridentata (Asteraceae) , 113 (AMNH). 28 mi SE of La Pine, June 25, 1979, R. T. Schuh, Cercocarpus ledifolius (Asteraceae) , 13 (AMNH). 4 mi NW of Worden on road to Keno, July 17, 1985, G. M. Stonedahl and J. D. McIver, Artemisia tridentata (Asteraceae) , 143 (AMNH). 5 mi N of Fort Klamath, 4400 ft, August 11, 1933, H. A. Scullen, 13 (USNM). 5–15 mi SE of Chiloquin, July 4, 1982, T. J. Henry and G. M. Stonedahl, Artemisia sp. (Asteraceae) , 13, 6♀ (USNM). Hayden Mt. Summit W of Keno, June 27, 1979, R. T. and Joe Schuh, 23 (AMNH). Lower Klamath Lake Hill, July 19, 1953, Joe Schuh, 33 (OSU). Lake Co.: 12 mi SE of Hampton, Glass Butte, July 12, 1968, Oman, 23 (OSU). E side Abert Lake, September 25, 1971, C Musgrave, Artemisia sp. (Asteraceae) , 13 (OSU). Hart Mountain, 6250 ft, September 2, 1977, J. Schuh, Chrysothamnus sp. (Asteraceae) , 23 (AMNH). Lost Forest, July 9, 1968, Oman, 13 (OSU). Umatilla Co.: 2 mi W of Tollgate, August 4, 1986, Schuh, Schwartz, Stonedahl, Holodiscus discolor (Rosaceae) , 13 (AMNH). Wallowa Co.: 30 mi N of Enterprise, July 30, 1968, Oman, 13 (OSU). Joseph, June 19, 1968 – July 18, 1975, Paul Oman, WF Chamberlain, 53 (OSU, TAMU). Wasco Co.: 24 mi SW The Dalles, Knebal Springs, 3900 ft, July 8, 1978, Nancy Herman, 13 (AMNH). Wheeler Co.: 2 mi W of Mitchell on Hwy 26, June 22, 1979, G. Stonedahl, Artemisia tridentata (Asteraceae) , 13 (AMNH). South Dakota: Brookings Co.: White , July 26, 1922, H. C. Severin, 13 (CAS). Custer Co.: Custer, Black Hills, August 26, 1935, M. W. Sanderson, 23 (KU). Black Hills, June 29, 1973, L.A. Kelton, 13 (CNC). Custer State Park, Black Hills National Forest, August 7, 1971, L.A. Kelton, 53, 15♀ (CNC). Fall River Co.: Hot Springs, August 27, 1922, H. C. Severin, 13 (CAS). Harding Co.: Buffalo, June 19, 1925, H. C. Severin, 1♀ (USNM). Wallworth Co.: Selby, September 25, 1925, H. C. Severin, 3♀ (USNM). Texas: Jeff Davis Co.: 39.7 km NW Ft. Davis, April 10, 1988, R. Anderson, 63 (TAMU). Utah: Box El­ der Co.: Hansel Valley, June 20, 1969, H. J. Hanson, 23 (USU). Kelton, June 27, 1969 – July 3, 1969, G. F. Knowlton, 63 (USU). Mantua, August 24, 1967, G. F. Knowlton, 43 (OSU). Cache Co.: 0.4 mi off Rt 189 along Franklin Basin Road, 6575 ft, July 18, 2001, T. J. Henry and A. G. Wheeler, Jr., Artemisia tridentata (Asteraceae) , 13, 3♀ (USNM). Beaver Mountain, August 17, 1982, G. F. Knowlton, 13 (USU). Blacksmith Fork Canyon, June 28, 1971, Ritcher and Carlson, G. F. Knowlton, 23 (OSU). Elk Valley, July 29, 1976, W. J. Hanson, 13 (USU). Green Canyon, June 22, 1955 – August 2, 1973, G. F. Knowlton, G. L Nielson, 23 (USU). Logan Canyon, July 29, 1934 – August 29, 1980, G. F. Knowlton, T. O Thatcher, 53 (USU). Providence, August 3, 1934 – June 2, 1953, J. Plant, C. Smith, 23 (USU). Daggett Co.: Red Springs Camp, Ashley National Forest, August 25, 1966, C. W. O’Brien, 43 (UCB). Davis Co.: Bountiful, July 2, 1935, G. F. Knowlton, 13 (USU). Farmington Canyon, July 3, 1935 – June 6, 1979, G. E. Bohart, G. F. Knowlton, 63 (USU). Duchesne Co.: 23.7 mi S of Myton, Wells Draw, T10S, R15E, 6000 ft, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 33 (AMNH). Garfield Co.: 10 mi S of Antimony, July 18, 1969, 13 (BYU). 14.3 mi S Rt 95 on Rt 276, 5000 ft, June 18, 1983, R. T. Schuh, M. D. Schwartz, 23 (AMNH). Boulder Mt., June 13, 1969, D. W. Davis, G. F. Knowlton, 53 (USU). Mt. Ellen, 9900 ft, August 27, 1981, T. Griswold, 13 (USU). Hatch, August 11, 1972, L.A. Kelton, 8♀ (CNC). Panguitch, June 27, 1927 – June 29, 1942, G. F. Knowlton, 33 (USU). Grand Co.: Colorado River Canyon near Fischer Towers, 4000 ft, May 27, 1978, D. A. and J. T. Polhemus, 43 (JTP). Iron Co. : Cedar City, June 29, 1965, H. H. Knight, 13, 1♀ (USNM). Juab Co.: 17 mi E of Leamington, July 27, 1962, C. W. O’Brien, 13 (UCB). Goshute Indian Reservation, July 27, 1953, P. D. Ashlock, 13 (BYU). Millard Co.: Scipio, June 29, 1965, H. H. Knight, 33 (USNM). Rich Co.: Garden City, July 7, 1969, G. F. Knowlton, 13 (USU). Logan Canyon, Bear Lake Summit, August 22, 1998, M. D. Schwartz, 1♀ (CNC). Monte Cristo, September 10, 1967, K. Capelle, 13 (USU). Salt Lake City Co.: Salt Lake City, July 13, 1976, Knowlton, Crowe, 13 (USU). Salt Lake Co.: Magna, June 16, 1954, G. F. Knowlton, 13 (AMNH). Mouth of Little Cottonwood Canyon on Rt 210, 5800 ft, June 28, 1981, M. D. Schwartz, Artemisia ludoviciana (Asteraceae) , 103 (AMNH). San Juan Co.: 1 mi W of Clay Hills Crossing Road on Rt 263, T39S, R15E, 5100 ft, June 18, 1983, R. T. Schuh and M. D. Schwartz, Artemisia filifolia (Asteraceae) , 13 (AMNH). 27 mi E of Blanding at mp 101 on Rt 95 near Mule Tower Ruins, 6500 ft, June 12, 1982, M. D. Schwartz, Artemisia ludoviciana (Asteraceae) , 13 (AMNH). 3 mi W of Clay Hills Crossing Road on Rt 263, T39S, R15E, 5000 ft, June 18, 1983, Schuh, Schwartz, Artemisia filifolia (Asteraceae) , 53 (AMNH). Bridges National Monument, April 12, 1969, G. F. Knowlton, 13 (USU). Brush Basin Rim Rd., Co. rd. no. 227 0.5 E mp 116, 5700ft, June 12, 1982, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 283 (AMNH). Natural Bridges National Park, July 12, 1967, G. F. Knowlton, 33 (USU). White Canyon, Duckett Crossing , May 29, 1985, J. T. and D. A. Polhemus, Artemisia tridentata (Asteraceae) , 63 (JTP). Sanpete Co.: 13 mi E of Fairview on Utah Rt 31, T14S, R6E, 7500 ft, August 8, 1981, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 133 (AMNH). Sevier Co.: 11 mi W of Sevier on I­70, 6400 ft, July 15, 1980, G. Stonedahl, Chrysothamnus viscidiflorus (Asteraceae) , 33 (AMNH). 11 mi W of Sevier on Rt 4, 2000 m, July 15, 1980, R. T. Schuh, G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 103 (AMNH). 32 mi N of Hwy 24 and Hwy 72, 7560 ft, July 17, 1980, G. Stonedahl, Atriplex sp. (Chenopodiaceae) , 93 (AMNH). Fish Lake National Forest, R2E, T25S, sec. 24, 25, 8720 ft, August 16, 1929, July 16, 1980, G. M. Stonedahl, Artemisia sp. (Asteraceae) , 13 (AMNH). Richfield, August 15, 1929, 13 (USNM). Salt Gulch, 12 mi N Hwy 24 on Hwy 72, 8060 ft, July 17, 1980, G. Stonedahl, Artemisia sp. (Asteraceae) , 113 (AMNH). Summit Co.: 3 mi SE of Bear River, August 5, 1971 – August 12, 1971, W. J. Hanson, 13 (USU). Coalville, July 13, 1965, H.H. Knight, det. as Psallus artemisicola Knight by Knight, 1♀ (CNC). 4.5 mi E of Oakley, Weber Canyon Road, August 19, 1986, G. M. Stonedahl, Artemisia tridentata (Asteraceae) , 63 (AMNH). Kimball Junction, August 31, 1967, L.A. Kelton, 23 (CNC). Uintah Co.: 22 mi N of Vernal, August 25, 1966, C. W. O’Brien, 43 (UCB). Bonanza, September 18, 1976 – July 7, 1982, G. E. Bohart, 23 (BYU). Bonanza, White River Shale Project Trailers, T 9S, R24E, sec. 23, 5000 ft, June 1, 1981 – June 8, 1981, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 303 (AMNH). Kane Hollow on Red Cloud Loop, 8500 ft, September 7, 1982, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 413 (AMNH). Unknown Co.: Cedar Point, July 15, 1952, G. E. Bohart, G. F. Knowlton, 13 (USU). Knaab, August 6, 1954, G. F. Knowlton, D. W. Dayle, 13 (KU). Utah Co.: American Fork, June 28, 1965, H.H. Knight, det. as Europiella decolor by H. H. Knight, 23, 1♀ (CNC). American Fork, June 28, 1965, H. H. Knight, 103, 10♀ (USNM). Provo, June 4, 1965, A. M. Wescloh, 13 (BYU). Wasatch Co.: 30 mi SE of Kamas on Rt 34, Uintah National Forest, Wolf Creek Campground, T4S, R10W, sec. 7, 9000 ft, August 15, 1986, Schwartz, Stonedahl, Artemisia tridentata (Asteraceae) , 313 (AMNH). Daniels Canyon, August 8, 1964, G. F. Knowlton, 13 (UCU). Washington Co.: Leeda Canyon, July 28, 1965, W. J. Hanson, 13 (USU). St. George, June 30, 1965, H. H. Knight, 103, 10♀ (USNM). Zion National Park, August 1, 1929 – June 18, 1935, C. C. Searl, G. F. Knowlton, 23 (SDNM). Zion National Park, Lava Point Campground, 7000 ft, May 24, 1981, M. D. Schwartz, Artemisia tridentata (Asteraceae) , 13 (AMNH). Wayne Co.: Hanksville, Vasco Tanner, 23 (BYU). Washington: Benton Co.: Hanford Site ALE, T11N, R26E, sec. 34, 1200 ft, June 1, 1994, R. S. Zack, 23, 4♀ (WSU). Kittitas Co.: 5 mi S of Ellensburg, T17N, R18E, sec. 34, June 13, 1989, R. S. Zack, 13, 4♀ (WSU). Okanogan Co.: 0.5 mi S of Malott, June 2, 1966, W. Gagne and J. Haddock, 23 (UCB). Okanogan, July 6, 1966 – June 22, 1983, W. Gagne, J. Haddock, S. G. Cannings, 83 (UBC, UCB). Whitman Co.: 4 mi NE of Steptoe, Steptoe Butte, June 11, 1989, R. S. Zack, 23 (WSU). Colfax , July 10, 1925, C. L. Fox, 13 (CAS). Fallon, June 8, 1949, C. O. Esselbaugh, 13 (AMNH). Yakima Co.: 11 mi SW of Ellensburg, T16N, R17E, sec. 3, Wenas Road, June 13, 1989, R. S. Zack, 23 (WSU). Naches, July 7, 1935, R. H. Beamer, 33 (KU). Yakima, September 6, 1932, A. R. Rolfs, 13, 1♀ (USNM). Wyoming: Albany Co.: 9 mi Laramie, 8600 ft, August 16, 1968, Oman, 53 (OSU). Near Keystone, July 26, 1981, J. T. and D. A. Polhemus, 13 (JTP). Big Horn Co.: 21 mi W of Burgess Jct on Rt 14, 8500 ft, August 13, 1986, Schuh, Schwartz, Stonedahl, Artemisia tridentata (Asteraceae) , 23 (AMNH). 27 mi W of Burgess Jct on Rt 14, 7700 ft, August 12, 1986, Schuh, Schwartz, Stonedahl, Artemisia tridentata (Asteraceae) , 553 (AMNH). Carbon Co.: 11 mi S of Red Lodge, Parkside Campground, 7250 ft, August 12, 1986, Schwartz, Stonedahl, Artemisia tridentata (Asteraceae) , 983 (AMNH). Saratoga, July 29, 1931, H. H. Knight, 23, 1♀ (USNM). Fremont Co.: Wind River Mts., 2.5 mi SW Shoshone NF boundary on Rt 131, August 14, 1986, Schwartz, Stonedahl, Artemisia tridentata (Asteraceae) , 473 (AMNH). Lincoln Co.: 4 mi N of W. Frontier, June 26, 1966, W. Gagne, J. Haddock, 13 (UCB). Kemmerer, July 13, 1983 – August 24, 1983, Bohart, Meadows, 43 (BYU). Natrona Co. : 7 mi W of Waltman, August 12, 1962, J. G. and B. L. Rosen, 13 (AMNH). Park Co.: Yellowstone National Park, 5 mi S of Tower Falls, 7350 ft, September 17, 1980, J. D. Lattin, 43 (OSU). Sheridan Co.: Arvada, August 19, 1927, H. H. Knight, 13 (USNM). SW of Sheridan, August 16, 1979, J. T. Polhemus, 13 (JTP). Sweetwater Co.: 30 mi E of Rock Springs near Point of Rocks, July 4, 1968, H. H. Knight, 13, 1♀ (USNM). 10 mi E of Rock Springs, July 4, 1968, H.H. Knight, det. as Europiella decolor by H. H. Knight, 1♀ (CNC).

Europiella lattini , new species

Figures 2 , 4

DIAGNOSIS: Recognized by its relatively small size, strongly infuscate hind femora, dark areas on the hemelytra (fig. 2), and very large male genitalia (fig. 4), including pygophore and vesica, of the type found in E. artemisiae and E. decolor . Distinguished from E. artemisiae by its smaller size and shorter second antennal segment relative to the width of the head in males; easily distinguished from E. artemisiae and E. decolor by the smaller pygophore and vesica and the longer anterior vesical spine in those two species.

DESCRIPTION: Male: Relatively small species, total length 2.81–3.05, width pronotum 0.91–0.97. COLORATION (fig. 2): Head, pronotum, and scutellum generally mostly dark; hemelytra often dark with pale areas at base of corium, base of cuneus, and along costal margin; lateral angles of mesoscutum orange; all antennal segments black; labium generally pale, segment 4 strongly infuscate; venter of body generally black; coxae black; trochanters pale; fore­ and middle femora generally orange with a few dark spots, never strongly infuscate, hind femora much more heavily infuscate; tibiae pale with black spines with distinct black bases. SURFACE AND VESTITURE: Surface and vestiture as in generic diagnosis. STRUCTURE: Moderately elongate, nearly parallel­sided, corial margin only weakly convex. GENITALIA (fig. 4): Pygophore very large relative to total size of abdomen; vesica very large, posterior apical spine of vesica short, sharply curving, hook­shaped, just surpassing gonopore, anterior spine only slightly longer and weakly decurved; structure of vesica very similar to that of E. artemisiae and E. decolor , but anterior spine much shorter and entire vesica much larger than in those species.

Female: Coloration similar to that in male. Body form much more strongly ovoid than male. Total length 2.50–2.89, width pronotum 0.89–0.91.

ETYMOLOGY: Named in honor of John D. Lattin, who nurtured my interest in the Heteroptera and the North American Miridae while I was an undergraduate student at Oregon State University, and whose efforts in the field produced the first specimens of this species examined by me.

HOSTS: Recorded from Achillea sp. (Asteraceae) , Lupinus latifolius (Fabaceae) , and couch grass ( Poaceae ). These are certainly not all breeding records. Only further collecting will confirm the actual hosts for this species.

DISTRIBUTION: Primarily coastal mountains from Oregon to British Columbia.

DISCUSSION: The genital capsule is large relative to the size of the abdomen, and the aedeagus is also relatively large, which, upon careful examination, make this species easily distinguishable from the very similar appearing E. decolor .

SPECIMENS EXAMINED: Holotype: USA: Oregon: Benton Co.: Grass Mt., Summit Prairie, Lattin, VIII­14–1980, general sweeping; deposit­ ed in the American Museum of Natural History. Paratypes: CANADA: British Columbia: 10 mi S of Merritt, July 19, 1959, L. A. Kelton, 13, 4♀ (CNC). 24 km E of Cherryville, August 25, 1989, G.G.E. Scudder, 23 (UBC). 9 mi N of Clinton, July 26, 1988, G.G.E. Scudder, 23, 3♀ (UBC). 9.5 km N of Cawston, July 13, 1991, G.G.E. Scudder, 13 (UBC). Cariboo, Westwick Lake, August 1, 1959, G.G.E. Scudder, 13 (UBC). Cariboo, Westwick Lake, August 17, 1962, G.G.E. Scudder, 13 (UBC). Cariboo, Westwick Lake, June 23, 1961, G.G.E. Scudder, 23, 1♀ (UBC). Fairview, White Lake , July 4, 1995, J. Jarrett, 13, (UBC). Grand Forks, July 10, 1950, H. R. MacCarthy, 43, 1♀ (CNC). Gray Creek, July 22, 1959, L. A. Kelton, 13, 2♀ (CNC). Lillooet, Seton Lake, June 3, 1926, J. McDunnough, 13, (CNC). Manning Prov. Park, Allison Pass, August 12, 1970, L. A. Kelton, 33 (CNC). Oliver, 7 mi E of Indian Reservation, July 8, 1959, L. A. Kelton, 53, 3♀ (CNC). Oliver, Madden Lake, July 8, 1959, L. A. Kelton, 13, 1♀ (CNC). Oliver, McIntyre Creek, July 4, 1959, L. A. Kelton, 13 (CNC). Osoyoos, Anarchist Mt., July 13, 1970 – July 20, 1959, L. A. Kelton, 33 (CNC). Salmon Arm, June 11, 1961, G.G.E. Scudder, 13 (UBC). Vancouver Island, Kelsey Bay, July 13, 1979, G.G.E. Scudder, 13 (UBC). Vancouver, July 28, 1922, W. Downes, 13 (UBC). Venables Valley entrance, July 26, 1983, G.G.E. Scudder, 13 (UBC). White Lake, Kearns Creek , June 29, 1996, G.G.E. Scudder, 23, 2♀ (UBC). USA: Oregon: Benton Co.: Grass Mt. Summit Prairie, August 14, 1980, Lattin, 4♀ (OSU). Mary’s Peak below campground, 3500 ft, August 17, 1970, Achillea sp. (Asteraceae) , 63, 5♀ (AMNH, OSU). Mary’s Peak, 14 mi W Corvallis, September 19, 1976, no collector, 23 (AMNH). Mary’s Peak, 4000 ft, July 29, 1966 – July 31, 1966, W. Gagne and J. Haddock, 103, 2♀ (UCB, AMNH). Mary’s Peak, August 10, 1971, P. W. Oman, 13 (OSU). Mary’s Peak, July 1, 1982, T. J. Henry and J. D. Lattin, 163, 6♀ (USNM, AMNH). Mary’s Peak, meadow below campground, 3500 ft, August 23, 1988, A. Asquith and J. D. Lattin, Lupinus latifolius (Fabaceae) , 153, 12♀ (AMNH, OSU, USNM). Hood River Co.: Mt. Hood, June 25, 1975, Hanson, Knowlton, 13 (USU). Jackson Co.: Siskiyou Summit, Old Road, 1260 m, July 22, 1999, M. D. Schwartz, 13 (CNC).

Europiella morrisoni , new species

Figures 2 , 4

DIAGNOSIS: Recognized by the nearly entirely pale dorsum, the entirely black antennae, and the body in males being long, narrow, and parallel­sided (fig. 1). Sexual dimorphism pronounced, the females being ovoid, with the hemelytra more or less conforming to shape of the abdomen (fig. 1). Most similar in coloration and body form to E. pilosula , but differing in having a slightly more elongate body form and black antennae. Apical vesical spines in male short, subequal in length, most similar to those of carvalhoi and pilosula .

DESCRIPTION: Male: Long slender species, total length 3.30–3.61. COLORATION (fig. 2): Head, pronotum, scutellum (except basally sometimes orange), hemelytra, and labium pale. Venter of thorax and abdomen weakly infuscate. Legs generally pale, hind femur with a few black spots apically, all tibiae with strongly contrasting black spots at the bases of black spines. SURFACE AND VESTITURE: Surface and vestiture as in generic description. STRUCTURE: Hemelytra conspicuously elongate, parallel­sided, apex of abdomen reaching only to base of cuneus; cuneus elongate triangular. GENITALIA (fig. 4): Pygophore only moderately large relative to total size of abdomen; apical spines of vesica short, of approximately equal length, anterior spine weakly notched.

Female: Coloration as in male; body form ovoid , much shorter than in male. Total length 2.50–2.89, width pronotum 0.93.

ETYMOLOGY: Names for Harold Morrison, who collected the only known specimens of this species.

HOSTS: Artemisia sp. (Asteraceae) .

DISTRIBUTION: Imperial Valley of southern California .

DISCUSSION: This species is known only from the type locality and appears never to have been recollected.

SPECIMENS EXAMINED: Holotype: 3, From Sagebrush beyond Jacumba, San Diego Co., Cal., swept 6:30 A.M., June 11, 1915, Harold Morrison; deposited in the National Museum of Natural History, Washington, D.C. Paratypes: USA: California: San Diego Co.: beyond Jacumba, swept 6:30 AM, June 11, 1915, Harold Morrison, Artemisia sp. (Asteraceae) , 113, 3♀ (USNM, AMNH).

Figures

Fig. 1. Habitus views of Europiella spp.: angulata–decolor. See appendix 1 for localities.

Fig. 2. Habitus views of Europiella spp.: decolor–unipuncta. Habitus of Knightopiella albipubescens. See appendix 1 for localities.

Fig. 4. Male genitalia of Europiella spp.: decolor–pintoi.

Fig. 10. Scanning electron micrographs of Europiella decolor (3). A. Head and thorax, lateral view. B. Mesothoracic spiracle and metepisternal scent­efferent system, lateral view. C. Pretarsus, lateral view. D. Setae on hemelytra, detail of microstructure.