Craspedometopon basale ( MATSUMURA, 1915 )

Rozkošný, R. & Kovac, D., 2007, Palaearctic And Oriental Species Of Craspedometopon Kertész (Diptera, Stratiomyidae), Acta Zoologica Academiae Scientiarum Hungaricae 53 (3), pp. 203-218 : 206-209

Craspedometopon basale ( MATSUMURA, 1915)

( Figs 1–7 )

Beris basalis MATSUMURA, 1915: 45

Acanthinoides basalis MATSUMURA, 1916: 368 (Pl. XXI, Fig 29) (secondary homonym of Beris basalis MATSUMURA, 1919 )

Craspedometopon frontal ; ÔUCHI, 1940 (Japanese specimens) (incorrect subsequent spelling)

Craspedometopon ussuricum KRIVOSHEINA, 1973: 180 , syn. n.

Craspedometopon frontale ; NAGATOMI 1975: 372.

Type material: Syntypes of both sexes of Beris basalis MATSUMURA as well as Acanthinoides basalis MATSUMURA originate from Japan and should be deposited in HUS ( WOODLEY 2001). Unfortunately, we did not succeed in borrowing these syntypes from Hokkaido University. Craspedometopon ussuricum KRIVOSHEINA is based on a male holotype and two female paratypes from Russia (Primorskiy district, Suputinskiy Reserve and Kedrovaya Pad’ Reserve). Although we did not examine the type series, N. P. KRIVOSHEINA kindly compared our material from North Korea and a male from Japan with types of C. ussuricum and was of the opinion that all specimens were conspecific.

Material examined: North Korea, Ryongaksan Mts., 10 km W of Pyongyang, 14.v. 1988, 1 m, M. KOZÁNEK; Kungansan Mts., Okryn Valley, 21.v.1988, 1 f, M. SLOVÁK; Wonsan, Botanical garden, 28.v. 1988, 2 mm, M. KOZÁNEK, in FSMU and ZIB, all identified as C. ussuricum by KRIVOSHEINA. South Korea, Kangwondo Province, Nam District, Magog, along Hongcheon river, 70 m, 37°43.786´N, 127°34.589´E, 24.v.–12.vi.2004, 1 f in USNM and 1 f in MHC; North Jeolla Province, Muju Prefecture, Anseong District, Muryeong Mt., 30.v.–6.vi.1999, 1 f in MNHN; South Chungcheong Province, Nami District, Keum Mt., Pohyeonsa, N 36°03.494 N, 127°27.225 E, 27.iv–14.v.2005, 1 f, 1.–8-vi. 2005, 1 m, 16 ff; 8.–24.vi.2005, 4 ff; North Chungcheong Province, Okcheon Prefecture, Dongi District, Soesan, 36°16.594 N, 127°36.742 E, 150 m, 19–28.vi. 2004, 4 mm, 12 ff; Jiri Mt., South Gyeongsang Province, Hamyang Prefecture, Macheon District, Samjeong, 700 m, 35°20.930 N, 127°38.503 E, 4 ff; Munsusa, 400 m, 35°24.739 N, 127°43.818 E, 9.vii.–17.viii.2005, 2 ff; all P. TRIPOTIN, all in MHC. Japan, Tokyo environs, 1906, 1 m, J. HARMAND; Osaka, Mt. Minoo, July 1930, 1 f, C. TETANISHI; Kyoto, 10.iv.1954, 1 f, W. C. BEN- TINCK; Camp Fuji, 7.v. 1954, 1 m, L. W. TELLER; Tamba, Sasayama, 4.v.1955, 1 f, 28.v. 1958, 1 m; Osumi, Sata, 27.iv. 1962, 1 m, all A. NAGATOMI, all in USNM, det. as C. frontale KERTÉSZ by A. NAGATOMI. Kagoshima Pref., Cape-Sata, 30.iv. 1966, 1 m, A. TANAKA, in FSMU, det. as C. ussuricum by N. P. KRIVOSHEINA.

Diagnosis. The eyes are bare or very short haired and the antennae black in the males or black to brown in the females. The hind femora and tibiae are completely black and the hind tarsi contrastingly yellow in the males. The medial incision on the hind margin of the male genital capsule is narrow.

Redescription: m. Head ( Fig. 1 ) black and blackish haired, occiput, both frontal triangles, face and postocular area whitish grey tomentose. Eyes bare or very short haired, contiguous for a long distance. Upper angle of lower frons shining black. Supra-antennal whitish tomentose patches separated by a deep, black medial groove, greyish tomentum on face less conspicuous. Ocellar tubercle, face, genae and lower postocular area with erect black hairs. Antenna entirely black, basal antennal segments black haired, arista only 1.3 times as long as antenna. Proboscis pale brown, yellowish haired, palpus usually black, shorter than half length of labellum, its apical segment slightly shorter and stouter than basal segment. Maximum width of lower postocular area about length of both basal antennal segments combined.

Thorax subshining black and densely punctate, punctation finer and denser than in C. orientale sp. n. Extreme lateral ridges of postpronotal calli reddish brown, apical halves of scutellar spines yellow and bare. Scutum and scutellum covered by dense black pile that is much longer then pedicel and predominantly erect especially in presutural area but short and semi-appressed on rest of scutum. Pleura also densely black haired, only posterior part of anepimeron and greater part of meron bare, shining black. Area above halteres indistinctly greyish tomentose.

Wings tinged with brown and yellow, basal half extending to discal cell brown and apical half yellowish or almost hyaline. Stronger veins in basal half dark brown, stigma and veins in apical half yellow. Costal sections between R 4 and R 5 twice as long as that between R 2+3 and R 4. Squamae small, their marginal hairs chiefly brown, halteres dark brown, stem more yellow.

Legs as in C. frontale but pale parts more limited, black and yellowish brown, pile black on dark parts and chiefly brown on pale parts. Femora dark, only fore femur yellowish at tip. All tibiae dark but fore and mid tibiae yellowish at both ends. Hind femora and tibiae completely black. Tarsi yellowish brown but hind tarsi contrastingly ochre yellow. Longer and erect black hairs visible on posterior surface of fore and mid femur.

Abdomen subshining black and densely punctate as on thorax. Abdominal pile inconspicuous, short and mainly appressed, chiefly black on disc but predominantly whitish to yellow on posterior half and venter. Longer black and erect hairs distinct at anterior corners both dorsally and ventrally.

Male terminalia ( Figs 5–7 ): Epandrium ( Fig. 5 ) elongate, with relatively shallow, semicircular proximal emargination, proctiger slightly longer than broad at base, tapered in distal half, cerci relatively stout and short, barely half as long as proctiger. Genital capsule ( Fig. 7 ) with narrow and short medial incision at distal margin. Gonostylus (4) markedly tapered in apical half, submedial ridge forming inner basal bilobate process. Ventrobasal lobe of gonostylus much more slender than in C. frontale . Aedeagal complex ( Fig. 6 ) relatively long, tripartite distally, with flattened proximal part as in other species.

Length: body 4.7–6.7 mm, wing 5.3–7.4 mm (13 males).

f: Head ( Fig. 2 ) black, occiput, paired supra-antennal patches, face, gena and distinctly swollen lower postocular area greyish tomentose. Eyes bare, postocular rim narrow but distinct in dorsal view ( Fig. 3 ), shining black, about as broad as lateral ocellus. Frontal vitta shining black, frontal index (length between anterior ocellus and base of antennae to width in middle) about 3.2 and frons thus relatively widest of all three known species, with usual narrow ridges along eye margins and punctation being about as dense as in C. frontale but shining bare area above low frontal calli usually much more extensive. Tomentose patches above antennae separated by a longitudinal, deep and anteriorly tapered medial groove. Antennae brown, flagellar complex darker and dull, larger than in male. Arista at most slightly longer than antenna. Proboscis and palpi brown, reaching about 3/4 of labellum length, apical segment stouter and longer than in male, almost black. Head pile indistinct, chiefly short and pale but chiefly blackish on postocular area where longer on lower part.

Thorax as in male but postpronotal calli, narrow subnotopleural line and postalar calli usually reddish brown. Brownish to black thoracic pile generally short and appressed. Basal infumation of wings less contrasting than in male but distinct, halteres yellowish to dark brown, only stem yellow basally. Legs as in male, tarsi often more brownish dorsally. Pile on legs predominantly pale, hairs on posterior surface of fore and mid femur shorter than in male and whitish. Abdominal pile inconspicuous, short and appressed, predominantly pale but partly black on disc. Female terminalia yellow with brown apical segment of cerci, genital furca similar to that of C. frontale .

Length: body 5.2–7.6 mm, wing 5.7–7.2 mm (41 females).

Variation: The antennal flagellar complex may be paler than in the male, often even pale brown though both basal segments usually black. The halteres also usually paler, yellowish brown in some specimens.

Note. Undoubtedly the type material of both MATSUMURA’ s taxa originated from Japan: Beris basalis from “ Hokkaido or Honshu” and Acanthinoides basalis from “ Hokkaido and Honshu”. Also records by ÔUCHI (1940) under “ C. frontal ” based on three females with blackish brown antennae from Takao Mt. near Tokyo, Mitsumine Mt. and Ohyama near Tokyo belong to this species. KRIVOSHEINA (1973) described her C. ussuricum from the Russian Far East (southern Sikhote Alin, Suputinskiy and Kedrovaya Pad’ Nature Reserves) and NAGATOMI (1975) recorded 73 specimens from the following localities under “ C. frontale ”: Hokkaido: Sapporo; Jozankei. Honshu: Tamba, Sasayama; Mt. Kogane near Sasayama; Okayama City. Shikoku: Ehime Pref., Jôjusha. Kyushu: Fukuoka Pref., Hikosan and Inunakiyama; Hyuga, Mt.Osuzu; Satsuma, Kurinodake; Kagoshima City; Osumi, Takakuma and Sata. South-West Islands: Yakushima, Miyanoura.

Distribution ( Fig. 24 ): Eastern Russia ( KRIVOSHEINA 1973 as C. ussuricum ), North Korea (material examined), South Korea (material examined), Japan (Palaearctic part, MATSUMURA 1915 as Beris basalis, MATSUMURA 1916 as Acanthinoides basalis, ÔUCHI 1940 as C. frontal, NAGATOMI 1975 as C. frontale and material examined).

mm (male terminalia)

Figures

Figs 1–7. Diagnostic characters of Craspedometopon basale: 1 = male head in lateral view, 2–3 = female head in lateral and frontal view, 4 = gonostylus in dorsal view, 5 = dorsal part of male terminalia, 6 = aedeagal complex, 7 = genital capsule. Scales:0.5 mm (heads), 0.2 mm (gonostylus), 0.3

Fig. 24. Distribution of Craspedometopon basale