Hemiphileurus dechambrei Ratcliffe, 2003

Ratcliffe, Brett C., 2003, New Species of Hemiphileurus Kolbe from Honduras and Guatemala (Coleoptera: Scarabaeidae: Dynastinae: Phileurini), The Coleopterists Bulletin 57 (3), pp. 334-338 : 335-338

Hemiphileurus dechambrei Ratcliffe , new species

(Figs. 4–5)

Type Material. Holotype male labeled ‘‘ Guatemala: Chimaltenango, Actenango , VIII-5-1965, approx 1500 M.’’ Allotype labeled ‘‘ Yepocapa, Guatemala, III-VI. 45, H. Elishewitz. ’’ Six paratypes with the following data: as holotype (2 males); ‘‘ Yepocapa, Guatemala, III-VI. 45, H. Elishewitz’ ’ (1 male); ‘‘ Guatemala: San Marcos, La Fraternidad, 1900 M, X-1996, J. Monzon’ ’ (1 male); (a) ‘‘ Guatemala, A. Alfaró,’’ (b) ‘‘ Museo Nacional Costa Rica (printed), Guatemala? (written), Col. A. Alfaro? (written),’’ (c) ‘‘ Costa Rica, INBIO, CR1001 , 114474 (1 male); ‘‘ San Marcos, Guatemala, IX.1966 ’’ (1 male) .

336 Holotype, allotype, and one paratype deposited at the University of Nebraska State Museum. Remaining paratypes deposited at the Instituto Nacional de Biodiversidad ( Santo Domingo de Heredia, Costa Rica), U.S. National Museum (Washington, D.C., currently at University of Nebraska State Museum), Roger-Paul Dechambre collection (Paris, France), and Brett C. Ratcliffe collection (Lincoln, NE) .

Description of Holotype. Male. Length 19.6 mm; width 8.9 mm. Color black. Head: Surface nearly smooth. Vertex and frons with deep, suboval hollow. Frons with 2 stout horns; horns long (subequal in length to width between eyes), slightly recurved, acuminate. Clypeus triangular, apex acute and strongly reflexed; strong carina extending from clypeal apex to base of each horn, region between each carina elevated from plane of disc. Eyes small, interocular width equals 10.0 transverse eye diameters. Antenna with 10 segments, club subequal in length to segments 2–7. Mandibles arcuate on lateral edge, apices acute. Pronotum: Surface with large, deep punctures; punctures sparse either side of median furrow and on sides, moderate in density on disc between furrow and sides, and dense (confluent) in anterior angles. Median, longitudinal furrow shallow, broad, extending anteriorly from near base to just past middle of disc; surface within furrow with sparse, moderate to large punctures, some confluent. Base with complete marginal bead. Elytra: Rows of punctures in furrows; punctures moderate to large, ocellate-umblicate, separated by about 1 puncture diameter or less between large punctures or 2 diameters between smaller punctures. Intervals moderately convex, all subequal in height, with sparse micropunctures. Pygidium: Surface moderately densely punctate; punctures moderate to large, minutely setigerous, becoming slightly sparser apically. In lateral view, surface regularly convex. Legs: Foretibia tridentate. Posterior tibia at apex with strong, acute tooth on upper angle and with several minute serrations and short, broad spinules below upper angle. Apex of first tarsomere of posterior tarsus extended into long, acute spine. Venter: Prosternal process short, columnar, apex bluntly rounded and with transverse groove, posterior surface at base with weak prominence. Last sternite with several moderately large punctures along basal margin, sparsely micropunctate elsewhere. Parameres: Figures 4–5.

Allotype. Length 18.7 mm; width 8.7 mm. Color reddish brown. As holotype except in the following respects: Head: Surface with dense, moderately large punctures, punctures sparse to absent laterad of clypeal carina. Vertex and frons lacking hollow. Frons with 2 strong tubercles, not horns. Pronotum: Median furrow extends to center of disc only. Pygidium: Surface densely punctate and slightly depressed at base either side of middle. In lateral view, surface weakly convex in basal 2/3, strongly convex in apical third.

Paratypes. Males (6). Length 16.5–20.3 mm; width 8.0– 9.2 mm. As holotype except in the following respects: Head: Horns reduced to large tubercles in 2 specimens. Pronotum: Median furrow extends from near base to center of disc or just past center. Pygidium: Surface densely punctate (punctures virtually contiguous), with large punctures.

Distribution. Hemiphileurus dechambrei is known only from Guatemala.

Locality Records (8 specimens examined). GUATEMALA (8) CHIMAL- TENANGO (5): Acatenango, Yepocapa; SAN MARCOS (2): La Fraternidad, San Marcos; No data (1).

Temporal Distribution. March to June (2), August (3), September (1), November (1).

Diagnosis. This species most closely resembles H. cylindroides (Bates) and H. simplex (Prell) because of the pronotal and elytral sculpturing, presence of horns (as opposed to tubercles) in the male majors, and the mostly finely punctate last sternite. The only reliable way to distinguish these species is by the form of the male parameres (Figs. 4, 6–7). Females can be identified only by being collected in association with males.

Nomenclature. Hemiphileurus cylindroides sensu Endrödi (1978, 1985) is, in fact, the new species described here! Endrödi clearly illustrated the parameres of what he thought to be Bates’ H. cylindroides , and, had he examined the Bates syntypes at the British Museum, the Museum National d’Histoire Naturelle, or the Canadian Museum 338 of Nature, he would have realized that he had these two species’ characterizations reversed. Consequently, the illustrations of the parameres in his 1985 manual are erroneous: his picture of H. costaricensis (Fig. 2131) is really H. cylindroides , and his picture of H. cylindroides (Fig. 2132) is really the un-named species that is described here as H. dechambrei .

Lastly, most of the countries listed for H. cylindroides sensu Endrödi refer to those countries from whence the types came for Bates’ H. cylindroides . Consequently, H. dechambrei (¼ cylindroides sensu Endrödi ) is not known to occur in Costa Rica and Panama.

Etymology. I am pleased to name this species in honor of Roger-Paul Dechambre (Paris, France) for his many contributions to the study of Dynastinae and for his discovery that the parameres of the Bates syntypes of H. cylindroides at the Museum National d’Histoire Naturelle were not the same as those illustrated by Endrödi.