Ipsviciopsis elegans Tillyard, 1922

Lambkin, Kevin J., 2020, Revision of the Ipsviciidae of the Late Triassic of Queensland (Hemiptera Cicadomorpha: Scytinopteroidea), Zootaxa 4860 (4), pp. 503-520 : 513

Ipsviciopsis elegans Tillyard, 1922

( Figs 33–52 )

Ipsviciopsis elegans Tillyard, 1922: 464 , text-fig. 85, plate liii, fig. 35; Tillyard, 1923: 484; Handlirsch, 1939: 9, 16; Evans, 1956: 248–249, figs 30B, 30E; Evans, 1963: 17–21, figs 1(c), plate 1B; Carpenter, 1992: 232; Jell, 2004: 45, 2 unnumbered figs.

Ipsvicia magna Tillyard, 1922: 465 , text-fig. 86; Tillyard, 1923: 484; Handlirsch, 1939: 9, 16; Evans, 1956: 248; Evans, 1963: 17; Jell, 2004: 45. Syn. nov.

Material. Holotype and paratype tegmina GSQ178a, GSQ278a, holotype tegmen of Ipsviciopsis magna GSQ 93a, Denmark Hill. Fifty additional specimens: Dinmore: fragmentary body specimens: QMF60199 a/b–60201a/b, 60202, 60203; tegmina: QMF60204 a/b–60209a/b, QMF60210 – 60232, 60236a/b, 60237, ACCI.54, I.55a/b, I.56, I.57, I.58a/b, I.65a/b, I.66a/b, I.172; clavi: QMF60233 –60235.

Description. Tegmen 13.2–16.2 mm long (holotype 13.3), 3.3–4.3 mm wide (holotype 3.4), length/width 3.8– 4.1; 1A about as long as PCu+1A, m-cua far distal to point of fusion of PCu and 1A; apex highly variable in shape and venation ( Figs 40–52 ), with no two specimens the same; entirely darkly shaded.

Notes. Except for the apparent absence of dSc (? a preservation artefact), the tegmen of Ipsvicia langenbergensis , from the Late Triassic of Germany, at about four times longer than wide, and with simple tuberculate/punctate sculpture, a very broadly convex costal margin, a forwardly inclined costal fracture, and similar apical venation, more or less meets the diagnosis of Ipsviciopsis and is thus formally transferred as Ipsviciopsis langenbergensis (Barth, Ansorge et Brauckmann) comb. nov. It differs from the type species in having a very short PCu+1A, much shorter than 1A, and m-cua at about the same level as the point of fusion of PCu and 1A, rather than far distal to it.

Figures

FIGURES 33–35. Ipsviciopsis elegans Tillyard, tegmen: 33, QMF60204a; 34, composite of QMF60228 (costal fracture) and QMF60204a/b; 35, QMF60210.

FIGURES 36–39. 36–38, Ipsviciopsis elegans Tillyard, body specimens: 36, QMF60203; 37, QMF60202; 38, QMF60199a; 39, possible ipsviciid nymph, QMF60198.

FIGURES 40–52. Ipsviciopsis elegans Tillyard, tegmen apex (excluding marginal membrane), variation in shape and venation, all QMF except Fig. 48: 40, 60206b; 41, 60227; 42, 60212; 43, 60218; 44, 60216; 45, 60223; 46, 60208b; 47, 60217; 48, ACCI.56; 49, 60205a; 50, 60210; 51, 60220; 52, 60211.