Macrocheilus nigrotibialis Heller, 1900

Zhao, Danyang & Tian, Mingyi, 2012, Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines, ZooKeys 228, pp. 77-100 : 88

Macrocheilus nigrotibialis Heller, 1900 Figs 7, 15, 23, 33, 34, 42, 54

Macrochilus nigrotibialis Heller 1900: 2. Andrewes 1920: 497; Csiki 1932: 1574; Lorenz 2005: 512. Type locality: Indonesia (Sulawesi), deposited in MDSG.

Diagnosis.

Length 8.0-10.1 mm, width 3.0-3.5 mm. Labrum (Fig. 7) convex, arcuate at apex, front setae beneath apex, intermediate one on apical margin, hind one close to middle margin; mandibles sharp at apex; mentum (Fig. 15) setose at base; median tooth triangular and shorter than lobes, with two pairs of setae at base; lobes obtuse at apex; maxillary palpomere 4 (Fig. 23) roundly dilated, labial palpomere 4 triangular in shape and dilated, 3 not dilated and bisetose on inner sides; ligula thickened, apical margin arcuate inwards, deeply depressed at sides, with a median impression close to apex, outer apical angles rounded, with a seta on either of median impression close to apex. Elytra with front spot large, just before middle, almost rounded, covering intervals 3-7 and extended to small part of 2 and 8; hind spot on inner angles, more or less triangular, covering intervals 1-5.

Male genitalia. Median lobe slender, apical lamella round (Figs 33-34).

Female genitalia. Gonocoxite slender, arcuate, with three or four setae on dorsal side, apex slightly obtuse (Fig. 42).

Remarks.

This species differs from following two species in having the tibiae black, maxillary palpomere 4 more dilated, the median lobe of male genitalia stouter and the apical lamella more rounded.

Materials examined.

1 male, the holotype, “typus!”, "Drs. Sarasin N. Celebes Panot-Maimang", “12623”, "Staatl. Museum fur Tierkunde. Dresden", deposited in SNSD. 1 female, labeled "Nord Borneo Mont Kina Balu 5-8, 1903, John Waterstradt; Macrochilus nigrotibialis Heller, det. Andrewes", deposited in MNHN.

Distribution.

Indonesia (Sumatra, Malaysia and Sulawesi).