Cynoglossum teheranicum Riedl (1963: 512)

Sutorý, Karel, 2024, Contribution to Cynoglossum species (Boraginaceae) in Turkey, Phytotaxa 633 (3), pp. 241-252 : 244-246

Cynoglossum teheranicum Riedl (1963: 512)

Holotype:— IRAN. Persien: Zentral-Elburs: am Südabhang des Totschal im Tal Häfthous NW Teheran; Sand- u. Kiesboden des Bachbettes , 1300–1500 m, 4 July 1948, P. Aellen 1426 (Holotype W 1961-0001745 A!, https://w.jacq.org/ W19610001745 A).

Paratype:— IRAN. Persien: Zentral Elburs: Am Südabhang des Totschal im Tal von Pas Qaleh [Pas e Qualeh] nördlich von Teheran, Bergsteppe, 1300–1600 m, 8 July 1948, P. Aellen 1440 ( W 1961-0001745 B!, https://w.jacq.org/ W 19610001745 B).

Description:—Biennials. Stem erect, simple, rarely several, only sparsely branched in upper part, 35–45 cm tall. Trichomes smooth, crisply, ± patent or subadpressed. Subbasal leaves mostly dry in flowering time, linear-lanceolate, long petiolate, glabrous on upper side. Cauline leaves linear-lanceolate or linear-oblong, 15–70 mm long, 3–8 mm wide, acute or obtuse, lower ones petiolate, upper ones with attenuate base, superior leaves sessile, uninervate, margins sometimes undulate, trichomes basiscopic, 0.5 mm long on lower leaves, but acroscopic, longer, about 0.7 mm long, sparse, adpressed on upper leaves. Inflorescence terminal, sparse (up to 6 branches), rarely also with sporadic short axial branches. Branches simple or some lower ones rarely furcate, small lanceolate bracts usually singly on lower branches, fructiferous pedicels patent, 3–4 mm long. Calyx lobes 1.5–2 mm long, oblong, later up to 7 mm long, 1 mm wide, subacute or obtuse, inner side glabrous ( Figure 5 ), outer side with 0.5 mm long, stout, appressed trichomes. Corolla small, ca. 5 mm long ( Figure 3 ). Nutlets broadly ovate, (6.5–) 6.6–7.0 (–7.3) tall, L/W ratio (1.13–) 1.22– 1.31 (–1.37), disc (abaxial part of nutlets) subplane, regularly glochidiate, glochidia slim, 1–1.5 mm long, with small intermixed tubercles between glochidia. Style 3–4 mm long.

Distribution:—Distributed in a belt from northern Iran (Alborz) through southern Azerbaijan, Armenia, and Georgia? up to northeast and central Turkey ( Fig. 6). Its distribution range well coincides with the ranges of species in the group of “ Irano-Caucaso-Anatolian species ” as defined by Noroozi et al. (2007).

Examined specimens and specimens cited by Riedl (1967):— IRAN: Mazandaran: In faucibus calc. vallis Chalus inter Valiabad et Pol-e Zanguleh, 25 June 1974, K.H. Rechinger Iter Iranicum VIII. 48408 (E, G, K, W); Prov. Mazandaran: in valle fluvii Čalus, ca. 2200 m, Pol-e Zanguleh, 6–7 August1948, K.H. Rechinger, Iter Iranicum II 6344 (G, US, W, Riedl 1967); Prov. Mazandaran: in valle fluvii Chalus, ca. 2400 m, 9 June 1937, K.H Rechinger 967 ( US, W, Riedl 1967); Quazvin: Persia borealis: in valle Lur [Loura] (Elburs occid.), 2000 m, 21 March 1902, J. & A. Bornmüller Iter persicum alterum 7733 (B, Riedl 1967); Persia boreali: Elburs, Nordhang des Kandovan, 2400 m, 9.6.1937, D.E. Gauba Pl. Gaubaeanae Iranicae 1639 (B); Près de Siah-Bishé [Siah Bisheh], N Kandavan, bord de route, baisment humide, 2000 m, 7 June 1960, H. Pabot s.n. (G); Gilan: Bazar Luleman Buchse [Pir Bast-e Luleman?, Luleman Rural District—Gilan] ( Riedl 1967); Khosrava [Khosrowabad] prope Shahpur [Shähpür, Salmas] [1884], J.A. Knapp ( Riedl 1967). GEORGIA: — “Georg: Ziarat. Bge.ˮ [A.A. Bunge] Riedl (1967) located this collection in Iran, but the expression “Georg.” should indicate Georgia. Ziarat in Afghanistan (Golestan) cannot be excluded, but there is also a locality by the same name in southern Georgia [Aspindza Mta Didi Ziarat’i 1992 m]. AZERBAIJAN:—Talish. O. Kuntze ( Riedl 1967). ARMENIA: — Vayots Dzor Province, Yeghegis—meadow above river near the road ca. 1.5 km ESE of the village centre, 1753 m s.m., 9 June 2009, L. Ekrt, E. Ekrtová 4029 (BRNM); Megrinskij [Meghri] r-n [rayon]. Poljana v grab[ovo] -dubovom lesu nad pos[elkom] Vagravar [Vahravar], 1 June 1956, T.V. Egorova, N.N. Tzvelev, S.K. Czerpanov s.n. (LE). TURKEY: — A4 Ankara: Çubuk, Karagöl, 15 May 1974, S. Erik s.n. (HUB); A7 GümüŞhane: Zigana geçidi, Torul yolu, 1450 m, 4 June 1969, T. Baytop 15336 (ISTE); Armenia turcica: Szanschak Gümüschhane (Gümüsch: Sorda), 1894, P. Sintenis 5677 (B, JE, LE). A8 Erzurum: pendici detritico-pietrose lungo il torrente fra Ispir e el passo Ovitdag, ca. 1850 m, 18 June 2002, M. Bigazzi, F. Selvi 02.75 (FI). B5 Yozgat: Çekerek, Kabak tepe, NT [north] Silis in Korn, 1200–1300 m, 22 June 1980, R. Ilarslan s.n. (ANK). B6 Pontus australis: Yildiss Dagh in reg. alp., 2300 m, 7 June 1890, J. Bornmüller 2444 (B); Bozkir—Küçyksu üçpinar, 1800 m, 14 June 1968, R. Çetik, T. Ekim, E. Yurdakul s.n. (ANK). B7 Armenia turcica: Kainandagh, inter Egin/Kemaliye, et Arabkir [Arapkir], in saxosis, 24 June 1889, P. Sintenis 920 (LD). B8 NE Turkey, Betw. Mus and Erzurum, 26 km N of Mus, overgrazed steppe with forest remnants, calcareous soil, eastern exposure, 1300–1500 m, 5 July1963, M. Zohary 57147 (HUJ).

Figures

FIGURE 3. Identification characters on nutlets of C. montanum, C. anatolicum, C. teheranicum and C. longeglochidiatum.

FIGURE 5. Cynoglossum teheranicum, flower.